Euphorbia hypericifolia |
Euphorbia crenulata |
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graceful sandmat, graceful spurge |
Chinese caps, western wood spurge |
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Habit | Herbs, annual, with taproot. | Herbs, usually biennial, occasionally annual, with taproot. |
Stems | erect to ascending, 15–50 cm, glabrous. |
erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
Leaves | opposite; stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous; petiole 1–3 mm, glabrous; blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally. |
petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous; glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary; appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire. |
campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
Staminate flowers | (0–)2–20. |
11–18. |
Pistillate flowers | ovary glabrous; styles 0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
Capsules | depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous; columella 1–1.1 mm. |
subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
Seeds | with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges. |
cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
Cyathia | in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia; peduncle 0.5–1.8 mm. |
peduncle 0–0.5 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
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Euphorbia hypericifolia |
Euphorbia crenulata |
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Phenology | Flowering and fruiting early spring–late fall. | Flowering and fruiting spring–summer. |
Habitat | Open, disturbed areas, nurseries. | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. |
Elevation | 0–200 m. (0–700 ft.) | 30–1800 m. (100–5900 ft.) |
Distribution |
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
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CA; CO; NM; OR
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Discussion | Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 301. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce glomerifera, C. hypericifolia, E. glomerifera | Tithymalus crenulatus |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) |
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