Euphorbia hypericifolia |
Euphorbia astyla |
|
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graceful sandmat, graceful spurge |
alkali spurge, Pecos spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with woody or fibrous-fleshy taproot, 5–12 mm thick. |
Stems | erect to ascending, 15–50 cm, glabrous. |
decumbent, ascending, or erect, few to many emerging from woody crown, 5–25(–50) cm, glabrous. |
Leaves | opposite; stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous; petiole 1–3 mm, glabrous; blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally. |
opposite; stipules connate into deltate scale, 0.2–0.5 mm, minutely lacerate at apex, glabrous; petiole 0–0.2(–0.3) mm, glabrous; blade orbiculate-reniform to acute-cordate, 2–5(–8) × 2–5(–6) mm, base ± asymmetric, cordate to auriculate, sometimes clasping stem, margins entire, apex narrowly acute, surfaces glabrous; 2- or 3-veined from base, but usually only midvein conspicuous. |
Involucre | obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous; glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary; appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire. |
broadly campanulate, 0.8–1.4 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to brownish, oblong, 0.2–0.3 × 0.5–0.7 mm; appendages white, flabellate to oblong, 0.1–0.2(–0.5) × 0.4–0.8 mm, distal margin entire or dentate-crenate. |
Staminate flowers | (0–)2–20. |
22–26. |
Pistillate flowers | ovary glabrous; styles 0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.3–0.4 mm, unbranched, thickened-clavate. |
Capsules | depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous; columella 1–1.1 mm. |
ovoid and broadly triangular, 1.5–1.9(–2.5) × 1.4–1.6(–2.2) mm, glabrous; columella 1.2–1.8 mm. |
Seeds | with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges. |
white, oblong, 4-angled in cross section, adaxial faces slightly concave, with long raphe between, 1.5–1.8 × 0.7–1 mm, markedly foveolate, with irregular to ± parallel or anastomosing ridges. |
Cyathia | in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia; peduncle 0.5–1.8 mm. |
solitary at distal nodes; peduncle 0.3–1(–1.5) mm. |
2n | = 28. |
|
Euphorbia hypericifolia |
Euphorbia astyla |
|
Phenology | Flowering and fruiting early spring–late fall. | Flowering and fruiting late spring–early fall. |
Habitat | Open, disturbed areas, nurseries. | Desert, grasslands, limestone substrates, usually on very saline or alkaline soils. |
Elevation | 0–200 m. (0–700 ft.) | 700–1100 m. (2300–3600 ft.) |
Distribution |
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
|
TX; Mexico (Coahuila, Nuevo León) |
Discussion | Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia astyla is a specialist on halophytic, alkaline soils and is known in the flora area only in part of Pecos County. The species is closely related to E. jejuna but differs in its sessile or sub-sessile leaves with a cordate-auriculate base and involucral gland appendages that are not deeply lobed or cleft. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce glomerifera, C. hypericifolia, E. glomerifera | Chamaesyce astyla |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Engelmann ex Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862) |
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