Euphorbia humistrata |
Euphorbia simulans |
|
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spreading sandmat |
mimicking sandmat, similar spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, annual or short-lived perennial, with usually slender, occasionally slightly thickened, rootstock. |
Stems | prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). |
prostrate to reclining, 5–40 cm, glabrous. |
Leaves | opposite; stipules distinct, linear-subulate, often irregularly 2- or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. |
opposite; stipules distinct, subulate, 0.5–0.7 mm, glabrous; petiole 0.7–1.3 mm, glabrous; blade orbiculate, oval, to shortly oblong, 1–3.2 × 1.5–5 mm, base subsymmetric, rounded, margins entire, apex usually rounded, occasionally emarginate, surfaces glabrous; venation obscure, only midvein conspicuous. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
turbinate to campanulate, 0.8–1.2 × 0.7–1 mm, glabrous; glands 4, red to purple, slightly concave, elliptic, 0.2–0.3 × 0.4–0.5 mm; appendages absent. |
Staminate flowers | 2–5. |
15–36. |
Pistillate flowers | ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid nearly entire length. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. |
broadly ovoid, 1.3–1.8 × 1.5–2 mm, glabrous; columella 1.1–1.4 mm. |
Seeds | white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate. |
whitish, reddish brown beneath coat, oblong, 4-angled in cross section, 1.5–2 × 1.3–1.8 mm, with 5–7 faint transverse ridges or wrinkles. |
Cyathia | solitary at distal nodes; peduncle 0.1–0.6(–2) mm. |
solitary at distal nodes; peduncle 0.3–0.7 mm. |
2n | = 28. |
|
Euphorbia humistrata |
Euphorbia simulans |
|
Phenology | Flowering and fruiting spring–late summer. | Flowering and fruiting year-round. |
Habitat | Stream and river banks, gravel bars, floodplains, pond edges, disturbed fields, railroads, roadsides. | Desert scrub, mountains, hills, canyons, arroyos, flats, roadsides, clay, sandy, gravelly, and rocky soils. |
Elevation | 0–300 m. (0–1000 ft.) | 600–1300 m. (2000–4300 ft.) |
Distribution |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; TN; TX; VA; WV
|
TX; Mexico (Chihuahua, Coahuila) |
Discussion | Euphorbia humistrata is distributed throughout the Mississippi River valley and along other major river systems in the central and eastern United States. There are scattered reports of this species as a waif or as introduced farther north and/or east (for example, Minnesota, New York, North Carolina, Ontario, South Dakota, and Wisconsin), but the authors have not been able to verify these occurrences. Euphorbia humistrata is similar to E. maculata and is often confused with that species in herbaria. It can be distinguished from E. maculata by its tendency to root at the stem nodes, its longer styles, and its seeds that lack low transverse ridges and that are more bluntly angled. When growing side-by-side, E. humistrata has an overall less congested appearance and its cyathia are not as numerous or crowded as those of E. maculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia simulans, which in the flora area is known only from Brewster, Hudspeth, and Presidio counties, is difficult to distinguish in the field from the sympatric E. theriaca var. theriaca, because they are mainly distinguished by seed morphology. The latter has smaller seeds with (two or) three (or four) prominent transverse ridges, whereas E. simulans has larger seeds that are slightly wrinkled. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 289. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce humistrata | E. polycarpa var. simulans, Chamaesyce simulans |
Name authority | Engelmann: in A. Gray, Manual ed. 2, 386. (1856) | (L. C. Wheeler) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) |
Web links |