Euphorbia humistrata |
Euphorbia purpurea |
|
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spreading sandmat |
Darlington's glade spurge, glade or Darlington's glade spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with thick rootstock. |
Stems | prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). |
erect, unbranched, 70–100(–130) cm, glabrous. |
Leaves | opposite; stipules distinct, linear-subulate, often irregularly 2- or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. |
petiole 0–2 mm; blade lance-oblong to oblanceolate-oblong, 50–100 × 13–30 mm, base attenuate to cuneate, margins entire, apex usually acute, sometimes obtuse to rounded, minutely apiculate, abaxial surface glabrate to sparsely pilose, adaxial surface glabrous; venation pinnate, midvein prominent. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
narrowly campanulate, 2.1–3 × 3–4.2 mm, glabrous; glands 5, elliptic to slightly reniform, 1–1.2 × 1.5–2.1 mm; horns absent. |
Staminate flowers | 2–5. |
10–15. |
Pistillate flowers | ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. |
ovary glabrous; styles 3–3.5 mm, 2-fid. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. |
globose, 4.5–5.2 × 6–6.8 mm, 3-lobed; cocci rounded, verrucose, sometimes minutely so, glabrous; columella 4–4.8 mm. |
Seeds | white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate. |
mottled silver-brown, ovoid-globose, 3–4 × 2.5–3.5 mm, smooth; caruncle subconic, reniform, 0.8–1.1 × 1.4–1.6 mm. |
Cyathia | solitary at distal nodes; peduncle 0.1–0.6(–2) mm. |
peduncle 0–1 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–6, each unbranched or 1–2 times 2-branched, occasionally appearing pendent; pleiochasial bracts lance-ovate, shorter than distal leaves; dichasial bracts distinct, cordate-deltate to reniform, base subcordate, margins entire, apex rounded; axillary cymose branches 0–10. |
|
Euphorbia humistrata |
Euphorbia purpurea |
|
Phenology | Flowering and fruiting spring–late summer. | Flowering and fruiting spring–fall. |
Habitat | Stream and river banks, gravel bars, floodplains, pond edges, disturbed fields, railroads, roadsides. | Dry to moist forests and slopes, rock outcrops, swamps or seeps, especially over calcareous rocks. |
Elevation | 0–300 m. (0–1000 ft.) | 50–1100 m. (200–3600 ft.) |
Distribution |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; TN; TX; VA; WV
|
DE; MD; NC; NJ; OH; PA; VA; WV |
Discussion | Euphorbia humistrata is distributed throughout the Mississippi River valley and along other major river systems in the central and eastern United States. There are scattered reports of this species as a waif or as introduced farther north and/or east (for example, Minnesota, New York, North Carolina, Ontario, South Dakota, and Wisconsin), but the authors have not been able to verify these occurrences. Euphorbia humistrata is similar to E. maculata and is often confused with that species in herbaria. It can be distinguished from E. maculata by its tendency to root at the stem nodes, its longer styles, and its seeds that lack low transverse ridges and that are more bluntly angled. When growing side-by-side, E. humistrata has an overall less congested appearance and its cyathia are not as numerous or crowded as those of E. maculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia purpurea is primarily an eastern Appalachian forest species, but it also occurs in Adams, Highland, and Pike counties in southern Ohio. It is listed as endangered by Maryland, New Jersey, Ohio, and Pennsylvania, and is in the Center for Plant Conservation's National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 308. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce humistrata | Agaloma purpurea, Galarhoeus darlingtonii |
Name authority | Engelmann: in A. Gray, Manual ed. 2, 386. (1856) | (Rafinesque) Fernald: Rhodora 34: 25. (1932) |
Web links |