Euphorbia humistrata |
Euphorbia porteriana |
|
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spreading sandmat |
Porter's sandmat |
|
Habit | Herbs, annual, with taproot. | Herbs, usually annual, sometimes perennial, with slender to occasionally thickened taproot, 5 mm diam.. |
Stems | prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). |
erect to ascending, sometimes slightly woody at base, not mat-forming, 15–60 cm, usually glabrous, young branches rarely very sparsely short pubescent. |
Leaves | opposite; stipules distinct, linear-subulate, often irregularly 2- or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. |
opposite; stipules usually distinct, occasionally connate basally, triangular, entire or 2–4 parted, apically ciliate, 0.3–0.4 mm, usually glabrous, rarely very sparsely short pubescent; petiole 0.5–1.5 mm, usually glabrous, rarely very sparsely short pubescent; blade ovate, oval, elliptic, oblong-elliptic, or linear-elliptic, 4–12 × 1–7 mm, base asymmetric, rounded or subcordate, margins usually entire, rarely sparsely serrate, apex obtuse to acute, sometimes mucronate, surfaces abaxially often purple or red, adaxially green, usually glabrous, rarely very sparsely short pubescent; 3-veined from base, only midvein conspicuous. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
turbinate to campanulate, 0.7–1 × 0.8–1 mm, usually glabrous, rarely very sparsely short pubescent; glands 4, brown, slightly concave, elliptic-oblong, 0.1–0.2 × 0.3–0.6 mm; appendages white or pink to dark red, usually oblong or flabellate, occasionally rudimentary and forming narrow rim at edge of gland, (0–)0.1–0.2 × 0.4–1 mm, distal margin entire or crenulate. |
Staminate flowers | 2–5. |
8–14. |
Pistillate flowers | ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. |
ovary glabrous, subtended by triangular calyxlike structure; styles 0.4–0.6 mm, 2-fid 1/2 length. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. |
broadly ovoid, 1–1.5 × 1.7–2.1 mm, glabrous; columella 1–1.4 mm. |
Seeds | white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate. |
reddish brown, ovoid, 4-angled in cross section, 0.7–1 × 0.6–0.7 mm, smooth or obscurely wrinkled. |
Cyathia | solitary at distal nodes; peduncle 0.1–0.6(–2) mm. |
solitary at distal nodes; peduncle 0.5–1.1 mm. |
Euphorbia humistrata |
Euphorbia porteriana |
|
Phenology | Flowering and fruiting spring–late summer. | Flowering and fruiting year-round. |
Habitat | Stream and river banks, gravel bars, floodplains, pond edges, disturbed fields, railroads, roadsides. | Pine rocklands, coastal scrub, open hammocks. |
Elevation | 0–300 m. (0–1000 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; TN; TX; VA; WV
|
FL |
Discussion | Euphorbia humistrata is distributed throughout the Mississippi River valley and along other major river systems in the central and eastern United States. There are scattered reports of this species as a waif or as introduced farther north and/or east (for example, Minnesota, New York, North Carolina, Ontario, South Dakota, and Wisconsin), but the authors have not been able to verify these occurrences. Euphorbia humistrata is similar to E. maculata and is often confused with that species in herbaria. It can be distinguished from E. maculata by its tendency to root at the stem nodes, its longer styles, and its seeds that lack low transverse ridges and that are more bluntly angled. When growing side-by-side, E. humistrata has an overall less congested appearance and its cyathia are not as numerous or crowded as those of E. maculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia porteriana is found in Miami-Dade County and on Big Pine Key, Monroe County. This species is closely related to E. blodgettii, E. garberi, and E. serpens. The capsules of E. porteriana are almost always completely glabrous, but the authors have seen three specimens with just a few scattered hairs on the keels. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly hairy capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 285. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce humistrata | Chamaesyce porteriana, C. porteriana var. scoparia, C. scoparia |
Name authority | Engelmann: in A. Gray, Manual ed. 2, 386. (1856) | (Small) Oudejans: Phytologia 67: 48. (1989) |
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