Euphorbia humistrata |
Euphorbia pergamena |
|
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spreading sandmat |
rocklands spurge, Southern Florida sandmat |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). |
prostrate to ascending, occasionally mat-forming, terete to slightly flattened, 5–18 cm, glabrous on lower surface, strigillose to sericeous on upper surface. |
Leaves | opposite; stipules distinct, linear-subulate, often irregularly 2- or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. |
opposite; stipules distinct, subulate, better developed on lower side of stem, 0.4–1.3 mm, glabrous or pilose; petiole 0.3–0.5 mm, puberulent, sericeous, or strigillose; blade oblong to ovate, 4–7 × 2–4 mm, base asymmetric, hemicordate, larger side sometimes amplexicaulous, margins entire or serrulate, apex obtuse, surfaces puberulent, sericeous, or strigillose; 3-veined from base, lateral veins inconspicuous. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
campanulate, 1–1.3 × 1–1.1 mm, canescent; glands 4, pinkish, oblong or reniform, 0.2 × 0.4–0.6 mm; appendages white to pink, lunate, unequal, those of proximal glands oblique, 0.6–1 × 1.4–1.6 mm, those of distal glands symmetric, 0.3 × 0.8–0.9 mm, irregularly undulate to incised. |
Staminate flowers | 2–5. |
15. |
Pistillate flowers | ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. |
ovary puberulent, sericeous or strigillose; styles 0.5–1 mm, 2-fid at apex. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. |
oblate, well exserted from involucre at maturity, 1–1.3 × 1.3–1.8 mm, puberulent, sericeous or strigillose; columella 1 mm. |
Seeds | white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate. |
whitish to gray, ovoid, 4-angled in cross section, 0.7–0.8 × 0.5–0.6 mm, with 3–4 transverse sulci alternating with low transverse ridges. |
Cyathia | solitary at distal nodes; peduncle 0.1–0.6(–2) mm. |
solitary at distal nodes or at nodes of short, congested, axillary shoots; peduncle to 0.5 mm. |
Euphorbia humistrata |
Euphorbia pergamena |
|
Phenology | Flowering and fruiting spring–late summer. | Flowering and fruiting year-round. |
Habitat | Stream and river banks, gravel bars, floodplains, pond edges, disturbed fields, railroads, roadsides. | Crevices of limestone outcrops in pinelands and pine-palm woods. |
Elevation | 0–300 m. (0–1000 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; TN; TX; VA; WV
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FL; West Indies (Cuba, Hispaniola) |
Discussion | Euphorbia humistrata is distributed throughout the Mississippi River valley and along other major river systems in the central and eastern United States. There are scattered reports of this species as a waif or as introduced farther north and/or east (for example, Minnesota, New York, North Carolina, Ontario, South Dakota, and Wisconsin), but the authors have not been able to verify these occurrences. Euphorbia humistrata is similar to E. maculata and is often confused with that species in herbaria. It can be distinguished from E. maculata by its tendency to root at the stem nodes, its longer styles, and its seeds that lack low transverse ridges and that are more bluntly angled. When growing side-by-side, E. humistrata has an overall less congested appearance and its cyathia are not as numerous or crowded as those of E. maculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia pergamena is a Florida-listed threatened species known within the flora area from only Miami-Dade and Monroe counties in extreme southern Florida. The species is often included as a subspecies of E. adenoptera (for example, D. G. Burch 1965), but here A. Herndon (1993b) is followed and it is treated as a distinct species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 282. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce humistrata | Chamaesyce adenoptera subsp. pergamena, C. pergamena, E. adenoptera subsp. pergamena |
Name authority | Engelmann: in A. Gray, Manual ed. 2, 386. (1856) | Small: Bull. Torrey Bot. Club 25: 615. (1898) |
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