Euphorbia humistrata |
Euphorbia angusta |
|
---|---|---|
spreading sandmat |
Blackfoot sandmat, narrow-leaf spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). |
erect, 12–43 cm, uniformly strigose. |
Leaves | opposite; stipules distinct, linear-subulate, often irregularly 2- or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. |
opposite; stipules distinct, linear-subulate or nodiform to papilliform nodiform to papilliform stipules often reddish brown, 0.1–0.7 mm, strigose; petiole 0.3–1.2 mm, strigose; blade: proximal ovate to ovate-elliptic, distal linear to elliptic-linear, 7–41 × 2–5 mm, distal leaf blades more than 6 times as long as wide, base asymmetric, cuneate to rounded, margins entire, often involute on drying, apex acute, surfaces usually short strigose, occasionally glabrous adaxially; venation pinnate, only midvein conspicuous. |
Involucre | obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
turbinate to campanulate-turbinate, 1–1.5 × 1–1.4 mm, strigose; glands 4, green to yellow-green, concave, narrowly oblong, 0.2–0.4 × 0.4–0.7 mm; appendages white, flabellate, 0.5–1.1 × 0.3–0.5 mm, distal margin shallowly and irregularly toothed. |
Staminate flowers | 2–5. |
16–26. |
Pistillate flowers | ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. |
ovary strigose; styles 0.3–0.5 mm, 2-fid at apex to almost 1/2 length. |
Capsules | ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. |
broadly ovoid, 2–2.6 × 2.5–3 mm, strigose; columella 1.6–2.2 mm. |
Seeds | white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate. |
white, ovoid, 4-angled in cross section, 1.7–2.2 × 1.1–1.2 mm, transversely low-ridged or wrinkled. |
Cyathia | solitary at distal nodes; peduncle 0.1–0.6(–2) mm. |
solitary at distal nodes; peduncle 1.1–2.4 mm. |
2n | = 28. |
|
Euphorbia humistrata |
Euphorbia angusta |
|
Phenology | Flowering and fruiting spring–late summer. | Flowering and fruiting early spring–fall. |
Habitat | Stream and river banks, gravel bars, floodplains, pond edges, disturbed fields, railroads, roadsides. | Rocky limestone soils. |
Elevation | 0–300 m. (0–1000 ft.) | 400–1200 m. (1300–3900 ft.) |
Distribution |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; TN; TX; VA; WV
|
TX; Mexico (Coahuila) |
Discussion | Euphorbia humistrata is distributed throughout the Mississippi River valley and along other major river systems in the central and eastern United States. There are scattered reports of this species as a waif or as introduced farther north and/or east (for example, Minnesota, New York, North Carolina, Ontario, South Dakota, and Wisconsin), but the authors have not been able to verify these occurrences. Euphorbia humistrata is similar to E. maculata and is often confused with that species in herbaria. It can be distinguished from E. maculata by its tendency to root at the stem nodes, its longer styles, and its seeds that lack low transverse ridges and that are more bluntly angled. When growing side-by-side, E. humistrata has an overall less congested appearance and its cyathia are not as numerous or crowded as those of E. maculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia angusta, which in the flora area is known from the trans-Pecos region to the Edwards Plateau, is easily recognized by its erect habit, linear leaves, and relatively showy involucral gland appendages with toothed margins. The species is closely related to E. acuta and the Mexican endemic E. johnstonii Mayfield (M. H. Mayfield 1991); it is not only morphologically distinctive but is also the only species in sect. Anisophyllum with C3 photosynthesis (R. F. Sage et al. 2011; T. L. Sage et al. 2011; G. L. Webster 1975). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 272. | FNA vol. 12, p. 260. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce humistrata | Chamaesyce angusta |
Name authority | Engelmann: in A. Gray, Manual ed. 2, 386. (1856) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
Web links |