Euphorbia hirta |
Euphorbia perennans |
|
---|---|---|
pillpod sandmat, pillpod spurge |
perennial sandmat, Terlingua spurge |
|
Habit | Herbs, annual or perennial, with slender to thickened taproot. | Herbs, perennial, with strongly thickened, woody rootstock. |
Stems | usually erect to ascending, rarely prostrate or decumbent, 10–50(–75) cm, usually both strigillose and hirsute. |
erect, 7–45 cm, glabrous. |
Leaves | opposite; stipules usually distinct, rarely connate at base, undivided or divided into 2–4 narrowly deltate to linear-subulate segments, 0.5–1.8(–2.9) mm, pilose, often with light-colored, minute circular glands at base; petiole 1–3 mm, usually both strigillose and hirsute; blade ovate to rhombic, 7–43 × 3–18 mm, base strongly asymmetric, one side rounded or slightly cordate to truncate, the other cuneate to attenuate, margins serrulate to double serrulate, apex acute, surfaces often with red spot in center, usually strigose to hirtellous, rarely glabrescent; 3–5-veined from base. |
opposite; stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous; petiole 0.8–2 mm, glabrous; blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous. |
Involucre | obconic, 0.6–1.2 × 0.4–0.9 mm, strigillose; glands 4, greenish to pink, circular, oblong, or reniform, 0.1–0.2 × 0.1–0.2 mm; appendages white to pink, flabellate, subcircular, or transversely oblong, rarely absent, (0–)0.1–0.6 × (0–)0.1–0.7 mm, distal margin usually entire, rarely slightly lobed. |
broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous; glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm; appendages absent. |
Staminate flowers | 2–8. |
35–45. |
Pistillate flowers | ovary strigillose, often canescent when young; styles 0.2–0.6 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.7–0.9 mm, 2-fid nearly entire length. |
Capsules | subglobose to slightly oblate, 1–1.3 × 1.1–1.6 mm, strigillose; columella 0.7–1 mm. |
subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous; columella 2.2–2.7 mm. |
Seeds | brownish red to orange or pink, narrowly ovoid, 4-angled in cross section, 0.7–0.9 × 0.5–0.7 mm, usually rugulose or with 3–6 low transverse ridges, rarely nearly smooth. |
white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled. |
Cyathia | in dense, axillary and terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia, axillary glomerules either sessile or at tips of elongated, leafless stalks; peduncle 0.4–2.1 mm. |
solitary at distal nodes or at nodes of short, axillary branches; peduncle 1.8–3 mm. |
Euphorbia hirta |
Euphorbia perennans |
|
Phenology | Flowering and fruiting year-round. | Flowering and fruiting spring–fall. |
Habitat | Disturbed areas, roadsides, vacant lots, desert grasslands, mesquite woodlands, riparian forests with cottonwoods and willows, flood plain forests, pinelands, deciduous forests. | Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. |
Elevation | 0–1500 m. (0–4900 ft.) | 900–1200 m. (3000–3900 ft.) |
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MS; NC; NM; NY; SC; TX; Mexico; Central America; South America; West Indies; Bermuda [Introduced in North America; introduced also in Europe, Asia, Africa, Australia]
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TX; Mexico (Chihuahua) |
Discussion | Euphorbia hirta is a widespread weed that is distributed throughout tropical and subtropical regions of the world. It is probably native to at least central Mexico, and its native range possibly extends from South America to the southern United States. In many places within the flora area the species is certainly introduced, and it has been recorded as a waif from several states (for example, Maryland, Michigan, and Virginia). Euphorbia pilulifera Linnaeus, a rejected name that is a heterotypic synonym of the Asian E. parviflora Linnaeus, has been misapplied to North American material of E. hirta in the past. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011). Euphorbia perennans is known in the flora area only from Brewster County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 271. | FNA vol. 12, p. 282. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce gemella, C. hirta, E. gemella | Chamaesyce perennans |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) |
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