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pillpod sandmat, pillpod spurge

royal sandmat

Habit Herbs, annual or perennial, with slender to thickened taproot. Herbs, annual or short-lived perennial, with slender taproot to thickened and woody rootstock.
Stems

usually erect to ascending, rarely prostrate or decumbent, 10–50(–75) cm, usually both strigillose and hirsute.

prostrate, usually mat-forming, terete to slightly flattened, 40 cm, lower surface glabrous, upper surface strigillose, pilose or villous.

Leaves

opposite;

stipules usually distinct, rarely connate at base, undivided or divided into 2–4 narrowly deltate to linear-subulate segments, 0.5–1.8(–2.9) mm, pilose, often with light-colored, minute circular glands at base;

petiole 1–3 mm, usually both strigillose and hirsute;

blade ovate to rhombic, 7–43 × 3–18 mm, base strongly asymmetric, one side rounded or slightly cordate to truncate, the other cuneate to attenuate, margins serrulate to double serrulate, apex acute, surfaces often with red spot in center, usually strigose to hirtellous, rarely glabrescent; 3–5-veined from base.

opposite;

stipules distinct, entire or divided into 3–4 subulate to filiform segments, 0.8–2 mm, usually pilose, rarely glabrous;

petiole 0.5–1 mm, pilose to villous;

blade oblong, ovate or narrowly obovate, 3–10(–12) × 2–6 mm, base strongly asymmetric, hemicordate, margins serrulate, apex obtuse to subacute, surfaces glabrous or slightly pilose; 3-veined from base, often only midvein conspicuous.

Involucre

obconic, 0.6–1.2 × 0.4–0.9 mm, strigillose;

glands 4, greenish to pink, circular, oblong, or reniform, 0.1–0.2 × 0.1–0.2 mm;

appendages white to pink, flabellate, subcircular, or transversely oblong, rarely absent, (0–)0.1–0.6 × (0–)0.1–0.7 mm, distal margin usually entire, rarely slightly lobed.

narrowly turbinate, 1–1.2 × 0.4–0.7 mm, pilose;

glands 4, yellow to pink, unequal, proximal pair oblong or linear, 0.1 × 0.3–0.4(–0.6) mm, distal pair oblong or subcircular, 0.1 × 0.1–0.2 mm;

appendages pink to reddish, unequal, on proximal glands oblique, 0.4–0.8(–1) × 0.8–1.4(–2) mm, on distal glands symmetric, 0.2–0.5 × 0.2–0.5 mm, slightly undulate to slightly crenate.

Staminate flowers

2–8.

5–15.

Pistillate flowers

ovary strigillose, often canescent when young;

styles 0.2–0.6 mm, 2-fid 1/2 length.

ovary pilose to strigillose in parts, glabrous in other parts;

styles 0.8–1.3 mm, usually unbranched, rarely 2-fid at apex, filiform.

Capsules

subglobose to slightly oblate, 1–1.3 × 1.1–1.6 mm, strigillose;

columella 0.7–1 mm.

ovoid-triangular, 1.2–1.5 × 1–1.4 mm, pilose to strigillose in parts, glabrous in other parts;

columella 1–1.3 mm.

Seeds

brownish red to orange or pink, narrowly ovoid, 4-angled in cross section, 0.7–0.9 × 0.5–0.7 mm, usually rugulose or with 3–6 low transverse ridges, rarely nearly smooth.

brown to light gray, ovoid, 4-angled in cross section, 0.8–1 × 0.4–0.5 mm, with 4 or 5 deep transverse sulci alternating with low transverse ridges.

Cyathia

in dense, axillary and terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia, axillary glomerules either sessile or at tips of elongated, leafless stalks;

peduncle 0.4–2.1 mm.

usually in small cymose clusters on congested, axillary branches;

peduncle rudimentary or to 0.2 mm.

Euphorbia hirta

Euphorbia indivisa

Phenology Flowering and fruiting year-round. Flowering and fruiting summer–fall.
Habitat Disturbed areas, roadsides, vacant lots, desert grasslands, mesquite woodlands, riparian forests with cottonwoods and willows, flood plain forests, pinelands, deciduous forests. Grasslands, oak forests, oak-mesquite woodlands, oak-juniper communities, rarely entering desert scrub.
Elevation 0–1500 m. (0–4900 ft.) 1000–2000 m. (3300–6600 ft.)
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; LA; MS; NC; NM; NY; SC; TX; Mexico; Central America; South America; West Indies; Bermuda [Introduced in North America; introduced also in Europe, Asia, Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico
[WildflowerSearch map]
Discussion

Euphorbia hirta is a widespread weed that is distributed throughout tropical and subtropical regions of the world. It is probably native to at least central Mexico, and its native range possibly extends from South America to the southern United States. In many places within the flora area the species is certainly introduced, and it has been recorded as a waif from several states (for example, Maryland, Michigan, and Virginia). Euphorbia pilulifera Linnaeus, a rejected name that is a heterotypic synonym of the Asian E. parviflora Linnaeus, has been misapplied to North American material of E. hirta in the past.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia indivisa is characteristic of grasslands and oak woodlands from extreme western Texas to southeastern Arizona. The species is often treated as a synonym of E. dioeca Kunth, but the two species are readily separable on the basis of their seeds. The seeds of E. indivisa possess deep transverse sulci, whereas those of E. dioeca are merely rippled or with low transverse ridges. Euphorbia dioeca is a weedy species that occurs widely throughout tropical America but has yet to be encountered within the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 271. FNA vol. 12, p. 273.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce gemella, C. hirta, E. gemella E. dioeca var. indivisa, Chamaesyce indivisa
Name authority Linnaeus: Sp. Pl. 1: 454. (1753) (Engelmann) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935)
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