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pillpod sandmat, pillpod spurge

square-seed spurge, squareseed or Clark Mountain spurge

Habit Herbs, annual or perennial, with slender to thickened taproot. Herbs, annual, with slender taproot.
Stems

usually erect to ascending, rarely prostrate or decumbent, 10–50(–75) cm, usually both strigillose and hirsute.

erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous;

branches arcuate.

Leaves

opposite;

stipules usually distinct, rarely connate at base, undivided or divided into 2–4 narrowly deltate to linear-subulate segments, 0.5–1.8(–2.9) mm, pilose, often with light-colored, minute circular glands at base;

petiole 1–3 mm, usually both strigillose and hirsute;

blade ovate to rhombic, 7–43 × 3–18 mm, base strongly asymmetric, one side rounded or slightly cordate to truncate, the other cuneate to attenuate, margins serrulate to double serrulate, apex acute, surfaces often with red spot in center, usually strigose to hirtellous, rarely glabrescent; 3–5-veined from base.

opposite;

petiole 1–3 mm, often indistinct, glabrous or puberulent;

blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous;

midvein conspicuous.

Involucre

obconic, 0.6–1.2 × 0.4–0.9 mm, strigillose;

glands 4, greenish to pink, circular, oblong, or reniform, 0.1–0.2 × 0.1–0.2 mm;

appendages white to pink, flabellate, subcircular, or transversely oblong, rarely absent, (0–)0.1–0.6 × (0–)0.1–0.7 mm, distal margin usually entire, rarely slightly lobed.

turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent;

involucral lobes divided into several linear lobes;

glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous;

appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous.

Staminate flowers

2–8.

10–12.

Pistillate flowers

ovary strigillose, often canescent when young;

styles 0.2–0.6 mm, 2-fid 1/2 length.

ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length.

Capsules

subglobose to slightly oblate, 1–1.3 × 1.1–1.6 mm, strigillose;

columella 0.7–1 mm.

broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels);

columella 1.9–2.5 mm.

Seeds

brownish red to orange or pink, narrowly ovoid, 4-angled in cross section, 0.7–0.9 × 0.5–0.7 mm, usually rugulose or with 3–6 low transverse ridges, rarely nearly smooth.

white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges;

caruncle 0.1 × 0.2 mm.

Cyathia

in dense, axillary and terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia, axillary glomerules either sessile or at tips of elongated, leafless stalks;

peduncle 0.4–2.1 mm.

peduncle 1–1.9 mm.

Cyathial

arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched;

pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower;

dichasial bracts similar in shape to distal leaves but smaller or highly reduced.

Euphorbia hirta

Euphorbia exstipulata

Phenology Flowering and fruiting year-round. Flowering and fruiting summer–fall.
Habitat Disturbed areas, roadsides, vacant lots, desert grasslands, mesquite woodlands, riparian forests with cottonwoods and willows, flood plain forests, pinelands, deciduous forests. Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands.
Elevation 0–1500 m. (0–4900 ft.) 800–2000 m. (2600–6600 ft.)
Distribution
from FNA
AL; AR; AZ; CA; FL; GA; LA; MS; NC; NM; NY; SC; TX; Mexico; Central America; South America; West Indies; Bermuda [Introduced in North America; introduced also in Europe, Asia, Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia hirta is a widespread weed that is distributed throughout tropical and subtropical regions of the world. It is probably native to at least central Mexico, and its native range possibly extends from South America to the southern United States. In many places within the flora area the species is certainly introduced, and it has been recorded as a waif from several states (for example, Maryland, Michigan, and Virginia). Euphorbia pilulifera Linnaeus, a rejected name that is a heterotypic synonym of the Asian E. parviflora Linnaeus, has been misapplied to North American material of E. hirta in the past.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 271. FNA vol. 12, p. 322.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Poinsettia
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce gemella, C. hirta, E. gemella E. exstipulata var. lata
Name authority Linnaeus: Sp. Pl. 1: 454. (1753) Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859)
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