Euphorbia hirta |
Euphorbia chamaesula |
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pillpod sandmat, pillpod spurge |
mountain spurge |
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Habit | Herbs, annual or perennial, with slender to thickened taproot. | Herbs, perennial, with thick rootstock. |
Stems | usually erect to ascending, rarely prostrate or decumbent, 10–50(–75) cm, usually both strigillose and hirsute. |
erect, branched, 40–90 cm, glabrous. |
Leaves | opposite; stipules usually distinct, rarely connate at base, undivided or divided into 2–4 narrowly deltate to linear-subulate segments, 0.5–1.8(–2.9) mm, pilose, often with light-colored, minute circular glands at base; petiole 1–3 mm, usually both strigillose and hirsute; blade ovate to rhombic, 7–43 × 3–18 mm, base strongly asymmetric, one side rounded or slightly cordate to truncate, the other cuneate to attenuate, margins serrulate to double serrulate, apex acute, surfaces often with red spot in center, usually strigose to hirtellous, rarely glabrescent; 3–5-veined from base. |
petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
Involucre | obconic, 0.6–1.2 × 0.4–0.9 mm, strigillose; glands 4, greenish to pink, circular, oblong, or reniform, 0.1–0.2 × 0.1–0.2 mm; appendages white to pink, flabellate, subcircular, or transversely oblong, rarely absent, (0–)0.1–0.6 × (0–)0.1–0.7 mm, distal margin usually entire, rarely slightly lobed. |
turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
Staminate flowers | 2–8. |
8–12. |
Pistillate flowers | ovary strigillose, often canescent when young; styles 0.2–0.6 mm, 2-fid 1/2 length. |
ovary glabrous; styles 1–1.8 mm, 2-fid. |
Capsules | subglobose to slightly oblate, 1–1.3 × 1.1–1.6 mm, strigillose; columella 0.7–1 mm. |
depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
Seeds | brownish red to orange or pink, narrowly ovoid, 4-angled in cross section, 0.7–0.9 × 0.5–0.7 mm, usually rugulose or with 3–6 low transverse ridges, rarely nearly smooth. |
gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
Cyathia | in dense, axillary and terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia, axillary glomerules either sessile or at tips of elongated, leafless stalks; peduncle 0.4–2.1 mm. |
peduncle 1–3 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
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2n | = 26. |
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Euphorbia hirta |
Euphorbia chamaesula |
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Phenology | Flowering and fruiting year-round. | Flowering and fruiting spring–summer. |
Habitat | Disturbed areas, roadsides, vacant lots, desert grasslands, mesquite woodlands, riparian forests with cottonwoods and willows, flood plain forests, pinelands, deciduous forests. | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. |
Elevation | 0–1500 m. (0–4900 ft.) | 1700–2700 m. (5600–8900 ft.) |
Distribution |
AL; AR; AZ; CA; FL; GA; LA; MS; NC; NM; NY; SC; TX; Mexico; Central America; South America; West Indies; Bermuda [Introduced in North America; introduced also in Europe, Asia, Africa, Australia]
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AZ; NM; Mexico (Chihuahua, Sonora)
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Discussion | Euphorbia hirta is a widespread weed that is distributed throughout tropical and subtropical regions of the world. It is probably native to at least central Mexico, and its native range possibly extends from South America to the southern United States. In many places within the flora area the species is certainly introduced, and it has been recorded as a waif from several states (for example, Maryland, Michigan, and Virginia). Euphorbia pilulifera Linnaeus, a rejected name that is a heterotypic synonym of the Asian E. parviflora Linnaeus, has been misapplied to North American material of E. hirta in the past. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 271. | FNA vol. 12, p. 300. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce gemella, C. hirta, E. gemella | Tithymalus chamaesula |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Boissier: Cent. Euphorb., 38. (1860) |
Web links |