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six-angle spurge

Caliche sandmat

Habit Herbs, annual, with taproot. Herbs, perennial, with strongly thickened and lignified rootstock.
Stems

erect, unbranched or branched, 30–70(–100) cm, sparsely hispid, occasionally densely so at distal nodes.

prostrate, usually mat-forming, terete to slightly flattened and winged, to 30 cm, villous to pilose.

Leaves

opposite;

stipules (0–)0.1 mm;

petiole 1–4 mm, pilose;

blade linear-filiform, linear, or elliptic, 21–40 × 0.9–7.5 mm, base attenuate, margins entire, apex acute, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous;

venation obscure, only midvein conspicuous.

opposite;

stipules distinct or connate at base, 2-fid to laciniate into 2–5 linear to subulate divisions, 0.5–1(–2) mm, glabrous or villous;

petiole 0.5–2 mm, villous;

blade usually ovate to oblong, rarely suborbiculate, 5–13 × 4–8 mm, base asymmetric, hemicordate, margins serrulate, apex obtuse, surfaces villous; 3-veined from base, usually only midvein conspicuous.

Involucre

campanulate, 1–1.5 × (1–)1.5–1.8 mm, hispid;

glands 5, green to deep red, elliptic to reniform, 0.5 × 0.8–1 mm;

appendages white to green, tinged red, deltate to ovate, 0.7–1.7 × (0.9–)1.3–1.5 mm, entire.

turbinate, 0.9–1.4 × 0.7–1.1 mm, villous;

glands 4, pink, oblong to reniform, 0.1–0.2 × 0.3–0.4 mm;

appendages white to pink, flabellate, ovate, or oblong, 0.3–0.5 × 0.4–0.7 mm, distal margin entire, crenulate, or erose, often puberulent-ciliate.

Staminate flowers

15–30(–40).

8–12.

Pistillate flowers

ovary glabrous;

styles 0.7–1.1 mm, 2-fid nearly entire length.

ovary villous;

styles 0.2–0.4(–0.5) mm, 2-fid 1/2 length.

Capsules

subglobose to broadly ovoid, 4.7–6.5 × 4.9–6.5(–7.1) mm, glabrous;

columella 3.5–4.5 mm.

broadly ovoid, 1.5–1.9 × 1.3–1.7 mm, villous (uniformly so, but most pronounced toward base and along keels);

columella 1.2–1.4 mm.

Seeds

dark brown or dark gray, ovoid, 3.4 × 2.7 mm, rugose, whitish glaucous;

caruncle absent.

white to gray or pink, narrowly pyramidal-ovoid, 4-angled in cross section, 1.1–1.3 × 0.4–0.6 mm, rugulose with shallow depressions separated by inconspicuous transverse ridges.

Cyathia

solitary in leaf axils or in terminal cymes or dichasia;

peduncle 1–2.1 mm, strigillose.

solitary at distal nodes or at nodes of leafy, congested, axillary branches;

peduncle 1–1.5(–2.5) mm.

Euphorbia hexagona

Euphorbia velleriflora

Phenology Flowering and fruiting late summer–fall. Flowering and fruiting summer.
Habitat Sand prairies, other sandy soil habitats, stream banks, sand bars, damp places. Disturbed habitats, particularly roadsides, grasslands, live oak-thorn scrub.
Elevation 200–1300 m. (700–4300 ft.) 50–80 m. (200–300 ft.)
Distribution
from FNA
AR; CO; IA; IL; KS; MN; MO; MT; NE; NM; OK; SD; TX; WI; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico; Central America (Guatemala) [Introduced in North America]
[BONAP county map]
Discussion

Euphorbia hexagona is native to the central United States and is most common from southern South Dakota to Oklahoma and northern Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia velleriflora is native and widespread from northern Mexico to Guatemala. The species is apparently a recent introduction in southern Texas (W. R. Carr and M. H. Mayfield 1993), as the first records are from the 1990s. It is also expected in southern Arizona due to the presence of collections made a few kilometers south of the border in the Mexican state of Sonora. Euphorbia velleriflora is very similar to E. stictospora, and the two are sometimes confused. The two can be readily distinguished on the basis of their styles: unbranched in E. stictospora and 2-fid in E. velleriflora. Also, the involucral gland appendages of E. velleriflora are ciliate with short hairs, whereas those of E. stictospora are glabrous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 247. FNA vol. 12, p. 292.
Parent taxa Euphorbiaceae > Euphorbia > sect. Alectoroctonum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Anisophyllum velleriflorum, Chamaesyce velleriflora
Name authority Nuttall ex Sprengel: Syst. Veg. 3: 791. (1826) (Klotzsch & Garcke) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862)
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