Euphorbia hexagona |
Euphorbia serpillifolia |
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six-angle spurge |
thyme leaf spurge, thyme-leafed spurge, thymeleaf sandmat |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. | ||||
Stems | erect, unbranched or branched, 30–70(–100) cm, sparsely hispid, occasionally densely so at distal nodes. |
prostrate to ascending, often mat-forming, 7–35 cm, glabrous, pilose, or villous. |
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Leaves | opposite; stipules (0–)0.1 mm; petiole 1–4 mm, pilose; blade linear-filiform, linear, or elliptic, 21–40 × 0.9–7.5 mm, base attenuate, margins entire, apex acute, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; venation obscure, only midvein conspicuous. |
opposite; stipules distinct, divided nearly to base into 3–5 subulate to filiform segments, these sometimes 2-fid toward apex or laciniate, 0.7–2.1 mm, glabrous; petiole 0.5–1 mm, glabrous, villous, or pilose; blade ovate, oblong, elliptic, or obovate, 3–13 × 2–7 mm, base asymmetric, rounded to oblique, margins usually entire in proximal 1/2 and serrulate in distal 1/2, rarely serrulate nearly to base, apex obtuse or truncate, surfaces often with red spot in center, not papillate, glabrous, villous, or pilose; weakly 3-veined from base, usually only midvein conspicuous. |
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Involucre | campanulate, 1–1.5 × (1–)1.5–1.8 mm, hispid; glands 5, green to deep red, elliptic to reniform, 0.5 × 0.8–1 mm; appendages white to green, tinged red, deltate to ovate, 0.7–1.7 × (0.9–)1.3–1.5 mm, entire. |
obconic, 0.6–1.1 × 0.6–1 mm, glabrous, villous, or pilose; glands 4, yellow to pink, usually oblong to reniform, 0.1 × 0.2–0.3 mm; appendages white to pink, oblong or flabellate, rarely absent, (0–)0.1–0.2 × (0–)0.3–0.4 mm, distal margin entire or shallowly lobed. |
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Staminate flowers | 15–30(–40). |
5–20. |
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Pistillate flowers | ovary glabrous; styles 0.7–1.1 mm, 2-fid nearly entire length. |
ovary glabrous, villous, or pilose; styles 0.3–0.4 mm, 2-fid 1/2 length. |
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Capsules | subglobose to broadly ovoid, 4.7–6.5 × 4.9–6.5(–7.1) mm, glabrous; columella 3.5–4.5 mm. |
broadly ovoid to oblate, cocci not elongated nor terminating in empty portion, 1.4–1.9 × 1.5–2, glabrous, pilose, or villous; columella 1.2–1.6 mm. |
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Seeds | dark brown or dark gray, ovoid, 3.4 × 2.7 mm, rugose, whitish glaucous; caruncle absent. |
pink, light brown, or grayish, ovoid to narrowly ovoid, 4-angled in cross section, 1–1.4 × 0.6–0.9 mm, smooth to dimpled or rugose, or with faint transverse ridges that do not interrupt abaxial keel. |
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Cyathia | solitary in leaf axils or in terminal cymes or dichasia; peduncle 1–2.1 mm, strigillose. |
solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 0.4–1.7 mm. |
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Euphorbia hexagona |
Euphorbia serpillifolia |
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Phenology | Flowering and fruiting late summer–fall. | |||||
Habitat | Sand prairies, other sandy soil habitats, stream banks, sand bars, damp places. | |||||
Elevation | 200–1300 m. (700–4300 ft.) | |||||
Distribution |
AR; CO; IA; IL; KS; MN; MO; MT; NE; NM; OK; SD; TX; WI; WY
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AZ; CA; CO; FL; GA; IA; ID; IL; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; NB; ON; QC; SK; Mexico; South America |
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Discussion | Euphorbia hexagona is native to the central United States and is most common from southern South Dakota to Oklahoma and northern Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Euphorbia serpillifolia is variable, especially in regard to seed sculpturing. L. C. Wheeler (1941) documented and discussed this variation, suggesting that within the United States and Canada the taxon could be further divided into various taxa. However, Wheeler refrained from actually proposing names and commented that further study was needed. The authors concur with Wheeler and maintain a broad delimitation of the species, pending a detailed study of variation throughout its range. Otherwise indistinguishable hairy plants are treated as subsp. hirtula. In contrast to the widespread typical subspecies, subsp. hirtula ranges from northern Baja California, Mexico, to central California. In this region the two subspecies sometimes grow together. The spelling of the specific epithet follows the original publication of the name and contrasts with the often-used variant serpyllifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 287. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce serpillifolia | |||||
Name authority | Nuttall ex Sprengel: Syst. Veg. 3: 791. (1826) | Persoon: Syn. Pl. 2: 14. (1806) | ||||
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