Euphorbia hexagona |
Euphorbia maculata |
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six-angle spurge |
euphorbe maculée, milk purslane, milk spurge, prostrate spurge, sandmat, spotted sandmat, spotted sandmat or spurge, spotted spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | erect, unbranched or branched, 30–70(–100) cm, sparsely hispid, occasionally densely so at distal nodes. |
usually prostrate, occasionally with ascending tips, often mat-forming, not rooting at nodes, 5–45 cm, densely and evenly short-sericeous to sericeous or villous. |
Leaves | opposite; stipules (0–)0.1 mm; petiole 1–4 mm, pilose; blade linear-filiform, linear, or elliptic, 21–40 × 0.9–7.5 mm, base attenuate, margins entire, apex acute, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; venation obscure, only midvein conspicuous. |
opposite; stipules distinct, linear-subulate, sometimes irregularly 2–3-lobed, 1–1.3 mm, sparsely short-sericeous to sericeous or villous; petiole 0.5–1.5 mm, moderately short-sericeous to sericeous or villous; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side usually angled and other ± truncate and expanded into small, rounded auricle, margins serrulate (longer side) or subentire (shorter side), apex rounded or broadly acute, abaxial surface pale grayish green, moderately to densely lanulose to villous, adaxial surface usually with irregular reddish streak along midvein, glabrate or with sparse, long, slender hairs; palmately veined at base, pinnate distally. |
Involucre | campanulate, 1–1.5 × (1–)1.5–1.8 mm, hispid; glands 5, green to deep red, elliptic to reniform, 0.5 × 0.8–1 mm; appendages white to green, tinged red, deltate to ovate, 0.7–1.7 × (0.9–)1.3–1.5 mm, entire. |
obconic, 0.8–1 × 0.6–0.8 mm, sparsely strigose to short-sericeous; glands 4, green to yellow-green, turning pink with age, usually ± unequal, narrowly oblong to nearly linear, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate to oblong, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
Staminate flowers | 15–30(–40). |
2–5. |
Pistillate flowers | ovary glabrous; styles 0.7–1.1 mm, 2-fid nearly entire length. |
ovary sericeous; styles 0.3–0.4 mm, 2-fid at apex. |
Capsules | subglobose to broadly ovoid, 4.7–6.5 × 4.9–6.5(–7.1) mm, glabrous; columella 3.5–4.5 mm. |
ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.4 mm, sparsely to moderately and evenly sericeous; columella 1–1.2 mm. |
Seeds | dark brown or dark gray, ovoid, 3.4 × 2.7 mm, rugose, whitish glaucous; caruncle absent. |
white to light brown, oblong-ovoid, sharply angular in cross section, 1–1.2 × 0.6–0.9 mm, with 3–4 low, transverse ridges that cross angles. |
Cyathia | solitary in leaf axils or in terminal cymes or dichasia; peduncle 1–2.1 mm, strigillose. |
solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 0.1–0.6 mm. |
2n | = 28. |
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Euphorbia hexagona |
Euphorbia maculata |
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Phenology | Flowering and fruiting late summer–fall. | Flowering and fruiting spring–fall. |
Habitat | Sand prairies, other sandy soil habitats, stream banks, sand bars, damp places. | Disturbed areas, fallow fields, gardens, sidewalk cracks, railroads, roadsides. |
Elevation | 200–1300 m. (700–4300 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AR; CO; IA; IL; KS; MN; MO; MT; NE; NM; OK; SD; TX; WI; WY
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AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; Mexico; Central America; South America; West Indies; Eurasia; Africa; Pacific Islands (New Zealand); Australia
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Discussion | Euphorbia hexagona is native to the central United States and is most common from southern South Dakota to Oklahoma and northern Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia maculata is a widespread weed in temperate latitudes, and it also occurs in cool climates at higher elevations in the tropics. It is presumed to be native to eastern and central North America, but given its extremely weedy tendencies, it is difficult to know for sure. It spreads readily in association with greenhouse plants and earth-moving activities, and it is notorious for its ability to colonize sidewalk cracks in the summer, even in congested cities. The name E. maculata was misapplied by most earlier botanists (for example, L. C. Wheeler 1941) to plants with ascending stems that are treated here as E. nutans. D. G. Burch (1966) reviewed the sources of data used by Linnaeus in his original description and concluded that the name E. maculata applies to this prostrate-stemmed taxon. For further discussion of the distinctions between E. maculata and the similar 47. E. humistrata, see the treatment of that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 275. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce maculata, C. mathewsii, C. supina, C. tracyi, E. supina | |
Name authority | Nuttall ex Sprengel: Syst. Veg. 3: 791. (1826) | Linnaeus: Sp. Pl. 1: 455. (1753) |
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