Euphorbia hexagona |
Euphorbia capitellata |
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six-angle spurge |
capitate sandmat, head sandmat, head spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual or perennial, with slender to thick and woody rootstock. |
Stems | erect, unbranched or branched, 30–70(–100) cm, sparsely hispid, occasionally densely so at distal nodes. |
usually ascending (but ranging from decumbent to erect), 15–50 cm, glabrous, strigillose, or pilose. |
Leaves | opposite; stipules (0–)0.1 mm; petiole 1–4 mm, pilose; blade linear-filiform, linear, or elliptic, 21–40 × 0.9–7.5 mm, base attenuate, margins entire, apex acute, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; venation obscure, only midvein conspicuous. |
opposite; stipules distinct, filiform or divided into 2–3 subulate-filiform segments, without dark, circular glands at base, 0.6–1.5 mm, pilose; petioles 0.6–1.3 mm, glabrous, pilose, or strigillose; blade ovate to narrowly ovate, 4–19 × 2–8 mm, base asymmetric, one side strongly cordate, other side rounded to slightly cordate, margins entire or serrulate (commonly nearly entire with few scattered teeth, often slightly thickened), apex acute, surfaces often with red spot in center, glabrous, pilose, or strigillose; weakly 3-veined from base, usually only midvein conspicuous. |
Involucre | campanulate, 1–1.5 × (1–)1.5–1.8 mm, hispid; glands 5, green to deep red, elliptic to reniform, 0.5 × 0.8–1 mm; appendages white to green, tinged red, deltate to ovate, 0.7–1.7 × (0.9–)1.3–1.5 mm, entire. |
narrowly obconic to narrowly campanulate, 0.8–1.6 × 0.7–1.3 mm, glabrous or pilose; glands 4, yellow-green to pink or maroon, circular to oblong, 0.2–0.4 × 0.2–0.5 mm; appendages white to light pink, oblong to reniform or flabellate, 0.2–1.1 × 0.5–1.7 mm, surfaces glabrous, distal margin entire. |
Staminate flowers | 15–30(–40). |
25–40. |
Pistillate flowers | ovary glabrous; styles 0.7–1.1 mm, 2-fid nearly entire length. |
ovary glabrous or pilose; styles 0.4–0.6 mm, 2-fid entire length. |
Capsules | subglobose to broadly ovoid, 4.7–6.5 × 4.9–6.5(–7.1) mm, glabrous; columella 3.5–4.5 mm. |
ovoid to oblate, 1.3–1.9 × 1.4–2.1 mm, glabrous or pilose; columella 1.1–1.7 mm. |
Seeds | dark brown or dark gray, ovoid, 3.4 × 2.7 mm, rugose, whitish glaucous; caruncle absent. |
pink to pinkish gray, narrowly ovoid to narrowly ovoid-oblong, 4-angled or weakly 3-angled in cross section, 0.9–1.5 × 0.5–0.7 mm, irregularly dimpled, sometimes also with faint transverse ridges that do not pass through abaxial keel. |
Cyathia | solitary in leaf axils or in terminal cymes or dichasia; peduncle 1–2.1 mm, strigillose. |
in dense, terminal capitate glomerules, with reduced, bractlike leaves subtending cyathia, at tips of main stems and short, leafy, axillary branches; peduncle 0.1–1.2 mm. |
2n | = 14. |
|
Euphorbia hexagona |
Euphorbia capitellata |
|
Phenology | Flowering and fruiting late summer–fall. | Flowering and fruiting year-round in response to sufficient moisture. |
Habitat | Sand prairies, other sandy soil habitats, stream banks, sand bars, damp places. | Gravelly washes, rocky slopes, basaltic talus, disturbed roadsides, primarily desert scrub, desert grasslands, riparian forests, rarely oak-juniper woodlands. |
Elevation | 200–1300 m. (700–4300 ft.) | 600–1600 m. (2000–5200 ft.) |
Distribution |
AR; CO; IA; IL; KS; MN; MO; MT; NE; NM; OK; SD; TX; WI; WY
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AZ; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Sinaloa, Sonora)
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Discussion | Euphorbia hexagona is native to the central United States and is most common from southern South Dakota to Oklahoma and northern Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia capitellata is a characteristic herb in the Sonoran Desert of southern Arizona, ranging east to extreme southwestern Texas. During peak flowering, plants are attractive due to the dense clusters of cyathia with well-developed involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 247. | FNA vol. 12, p. 262. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce capitellata, C. pycnanthema, E. pycnanthema | |
Name authority | Nuttall ex Sprengel: Syst. Veg. 3: 791. (1826) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) |
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