Euphorbia heterophylla |
Euphorbia simulans |
|
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Mexican fireplant, painted euphorbia |
mimicking sandmat, similar spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, annual or short-lived perennial, with usually slender, occasionally slightly thickened, rootstock. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
prostrate to reclining, 5–40 cm, glabrous. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
opposite; stipules distinct, subulate, 0.5–0.7 mm, glabrous; petiole 0.7–1.3 mm, glabrous; blade orbiculate, oval, to shortly oblong, 1–3.2 × 1.5–5 mm, base subsymmetric, rounded, margins entire, apex usually rounded, occasionally emarginate, surfaces glabrous; venation obscure, only midvein conspicuous. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
turbinate to campanulate, 0.8–1.2 × 0.7–1 mm, glabrous; glands 4, red to purple, slightly concave, elliptic, 0.2–0.3 × 0.4–0.5 mm; appendages absent. |
Staminate flowers | 8–15. |
15–36. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid nearly entire length. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
broadly ovoid, 1.3–1.8 × 1.5–2 mm, glabrous; columella 1.1–1.4 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
whitish, reddish brown beneath coat, oblong, 4-angled in cross section, 1.5–2 × 1.3–1.8 mm, with 5–7 faint transverse ridges or wrinkles. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
|
Cyathia | peduncle 0.9–1.5 mm. |
solitary at distal nodes; peduncle 0.3–0.7 mm. |
2n | = 28. |
= 28. |
Euphorbia heterophylla |
Euphorbia simulans |
|
Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting year-round. |
Habitat | Disturbed areas, roadsides. | Desert scrub, mountains, hills, canyons, arroyos, flats, roadsides, clay, sandy, gravelly, and rocky soils. |
Elevation | 0–500 m. (0–1600 ft.) | 600–1300 m. (2000–4300 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
|
TX; Mexico (Chihuahua, Coahuila) |
Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia simulans, which in the flora area is known only from Brewster, Hudspeth, and Presidio counties, is difficult to distinguish in the field from the sympatric E. theriaca var. theriaca, because they are mainly distinguished by seed morphology. The latter has smaller seeds with (two or) three (or four) prominent transverse ridges, whereas E. simulans has larger seeds that are slightly wrinkled. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 289. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | E. polycarpa var. simulans, Chamaesyce simulans |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | (L. C. Wheeler) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) |
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