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Mexican fireplant, painted euphorbia

Mojave spurge

Habit Herbs, annual, with taproot. Herbs, perennial, with thick rootstock.
Stems

erect-ascending, 20–100 cm, sparsely pilose to villous;

branches ± straight.

slender, ascending, often sinuous, many, densely branched near base, 10–50 cm, usually glabrous, usually glaucous.

Leaves

usually alternate, occasionally opposite proximally;

petiole 10–50 mm, pilose;

blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate;

venation pinnate, midvein prominent.

petiole 0–1 mm;

blade broadly oblanceolate to obovate, 10–20 × 3–9 mm, base usually acute, occasionally short-attenuate, rarely obtuse, margins entire, apex usually acute, occasionally obtuse, acuminate to cuspidate, surfaces usually glabrous, usually glaucous;

venation pinnate, sometimes obscure, midvein prominent.

Involucre

usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous;

involucral lobes divided into several linear, smooth lobes;

gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous;

appendages absent.

campanulate to broadly turbinate, 2.2–3 × 2–2.5 mm, glabrous;

glands 4, irregularly semicircular to trapezoidal or elliptic-truncate, 0.8–1.5 × 1–2.2, margins strongly crenate or dentate;

horns usually absent, if present then straight, 0.1–0.2 mm, generally equaling teeth on gland margin.

Staminate flowers

8–15.

12–20.

Pistillate flowers

ovary glabrous or puberulent;

styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length.

ovary glabrous;

styles 1–1.2 mm, 2-fid.

Capsules

broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent;

columella 2.1–2.8 mm.

oblong-ovoid, 3.5–4 × 3.8–5 mm, 3-lobed;

cocci rounded, smooth, glabrous;

columella 3.3–3.8 mm.

Seeds

brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows;

caruncle 0.1 mm.

gray to whitish, oblong cylindric, 2–3 × 1.5 mm, irregularly shallowly pitted to almost smooth;

caruncle conic, 0.6 × 0.6 mm.

Cyathial

arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern);

pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves;

dichasial bracts highly reduced, rarely absent in highly congested clusters.

arrangement: terminal pleiochasial branches (1–)3–4(–5), each 1–3 times 2-branched;

pleiochasial bracts broadly ovate to subcordate, usually similar in size to, occasionally wider than, distal leaves;

dichasial bracts distinct, broadly ovate to almost reniform, base obtuse, margins entire, apex obtuse, acuminate to cuspidate;

axillary cymose branches 0–2(–4).

Cyathia

peduncle 0.9–1.5 mm.

peduncle 0.3–1 mm.

2n

= 28.

Euphorbia heterophylla

Euphorbia schizoloba

Phenology Flowering and fruiting nearly year-round. Flowering and fruiting spring–summer.
Habitat Disturbed areas, roadsides. Desert mountains and canyon slopes, rocky and gravelly soils.
Elevation 0–500 m. (0–1600 ft.) 500–1800 m. (1600–5900 ft.)
Distribution
from USDA
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia schizoloba is a desert perennial that occurs on bluffs and ledges in the Mojave and Sonoran Deserts. Sparsely pubescent plants of E. schizoloba are known from Arizona and are best represented by several collections from the Mazatzal and Sierra Ancha mountains in Gila County from between 1000 to 1800 meters. George Engelmann published two names for this species almost simultaneously in 1861 (E. schizoloba and E. incisa). Although the authors have not been able to determine which publication has priority, Engelmann himself cited E. incisa as a synonym of E. schizoloba (in W. H. Brewer et al. 1876–1880, vol. 2), as did J. B. S. Norton (1899). Also, the type specimen at MO was annotated by Engelmann as E. schizoloba, and there is no mention of the name E. incisa on the sheet.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 323. FNA vol. 12, p. 309.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. geniculata, Poinsettia geniculata, P. heterophylla E. incisa, Tithymalus incisus, T. schizolobus
Name authority Linnaeus: Sp. Pl. 1: 453. (1753) Engelmann: Proc. Amer. Acad. Arts 5: 173. (1861)
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