Euphorbia heterophylla |
Euphorbia micromera |
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Mexican fireplant, painted euphorbia |
desert spurge, Sonoran sandmat, tiny sandmat |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
prostrate, mat-forming, 5–35 cm, glabrous or shortly pilose. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
opposite; stipules distinct, subulate, 0.2–0.4 mm, pilose; petiole 0.5–1.2 mm, glabrous or pilose; blade ovate to elliptic, 6–15 × 2–4 mm, base asymmetric, one side cuneate to rounded, other side rounded, margins entire, apex obtuse, surfaces glabrous or pilose; venation obscure or only midvein conspicuous. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
campanulate, 0.4–0.6 × 0.5–0.9 mm, glabrous or pilose; glands 4, red, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent. |
Staminate flowers | 8–15. |
2–5. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary usually glabrous, rarely pilose; styles 0.1–0.2 mm, 2-fid at apex. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
oblong, 1.3–1.5 × 1.1–1.3 mm, usually glabrous, rarely pilose; columella 1–1.2 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
light gray, narrowly ovoid, 4-angled in cross section, 0.9–1 × 0.5–0.6 mm, smooth to slightly rugose or with 1–4 faint transverse ridges that do not pass through abaxial keel. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
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Cyathia | peduncle 0.9–1.5 mm. |
solitary at distal nodes; peduncle 0.4–1.4 mm. |
2n | = 28. |
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Euphorbia heterophylla |
Euphorbia micromera |
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Phenology | Flowering and fruiting nearly year-round. | Flowering nearly year-round in response to sufficient rainfall. |
Habitat | Disturbed areas, roadsides. | Desert scrub, riparian woods with ash and willow, saltbush scrub, Joshua tree woodlands and grasslands, often in sandy or gravelly areas. |
Elevation | 0–500 m. (0–1600 ft.) | -20–1800 m. (-100–5900 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)
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Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 278. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | Chamaesyce micromera |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Boissier: Proc. Amer. Acad. Arts 5: 171. (1861) |
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