Euphorbia heterophylla |
Euphorbia hirta |
|
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Mexican fireplant, painted euphorbia |
pillpod sandmat, pillpod spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual or perennial, with slender to thickened taproot. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
usually erect to ascending, rarely prostrate or decumbent, 10–50(–75) cm, usually both strigillose and hirsute. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, rarely connate at base, undivided or divided into 2–4 narrowly deltate to linear-subulate segments, 0.5–1.8(–2.9) mm, pilose, often with light-colored, minute circular glands at base; petiole 1–3 mm, usually both strigillose and hirsute; blade ovate to rhombic, 7–43 × 3–18 mm, base strongly asymmetric, one side rounded or slightly cordate to truncate, the other cuneate to attenuate, margins serrulate to double serrulate, apex acute, surfaces often with red spot in center, usually strigose to hirtellous, rarely glabrescent; 3–5-veined from base. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
obconic, 0.6–1.2 × 0.4–0.9 mm, strigillose; glands 4, greenish to pink, circular, oblong, or reniform, 0.1–0.2 × 0.1–0.2 mm; appendages white to pink, flabellate, subcircular, or transversely oblong, rarely absent, (0–)0.1–0.6 × (0–)0.1–0.7 mm, distal margin usually entire, rarely slightly lobed. |
Staminate flowers | 8–15. |
2–8. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary strigillose, often canescent when young; styles 0.2–0.6 mm, 2-fid 1/2 length. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
subglobose to slightly oblate, 1–1.3 × 1.1–1.6 mm, strigillose; columella 0.7–1 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
brownish red to orange or pink, narrowly ovoid, 4-angled in cross section, 0.7–0.9 × 0.5–0.7 mm, usually rugulose or with 3–6 low transverse ridges, rarely nearly smooth. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
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Cyathia | peduncle 0.9–1.5 mm. |
in dense, axillary and terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia, axillary glomerules either sessile or at tips of elongated, leafless stalks; peduncle 0.4–2.1 mm. |
2n | = 28. |
|
Euphorbia heterophylla |
Euphorbia hirta |
|
Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting year-round. |
Habitat | Disturbed areas, roadsides. | Disturbed areas, roadsides, vacant lots, desert grasslands, mesquite woodlands, riparian forests with cottonwoods and willows, flood plain forests, pinelands, deciduous forests. |
Elevation | 0–500 m. (0–1600 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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AL; AR; AZ; CA; FL; GA; LA; MS; NC; NM; NY; SC; TX; Mexico; Central America; South America; West Indies; Bermuda [Introduced in North America; introduced also in Europe, Asia, Africa, Australia]
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Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia hirta is a widespread weed that is distributed throughout tropical and subtropical regions of the world. It is probably native to at least central Mexico, and its native range possibly extends from South America to the southern United States. In many places within the flora area the species is certainly introduced, and it has been recorded as a waif from several states (for example, Maryland, Michigan, and Virginia). Euphorbia pilulifera Linnaeus, a rejected name that is a heterotypic synonym of the Asian E. parviflora Linnaeus, has been misapplied to North American material of E. hirta in the past. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 271. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | Chamaesyce gemella, C. hirta, E. gemella |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Linnaeus: Sp. Pl. 1: 454. (1753) |
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