Euphorbia heterophylla |
Euphorbia cordifolia |
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Mexican fireplant, painted euphorbia |
heartleaf sandmat |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
prostrate, occasionally mat-forming, 10–43 cm, glabrous. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate at base, filiform, 1–1.2(–2.8) mm, usually glabrous, rarely pilose; petiole 0.4–1 mm, usually glabrous; blade ovate to oblong, 4.4–12 × 2.6–7.6 mm, base asymmetric, cordate to rounded, margins entire, apex rounded to mucronulate, surfaces glabrous; only midvein conspicuous. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
campanulate, 1–1.3 × 1–1.3 mm, glabrous; glands 4, yellowish to pink, elliptic, 0.3–0.5 × 0.5–1 mm; appendages whitish to pink, sometimes drying red, elliptic to ovoid, 1.1–1.5 × 1.2–1.9 mm, distal margin entire, retuse, or erose. |
Staminate flowers | 8–15. |
5–40. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.5–0.8 mm, 2-fid nearly entire length. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
ovoid, 2–3 mm diam., glabrous; columella 1.2–2.7 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
gray or tan with dark brown mottling, ovoid, bluntly 3–4-angled in cross section, 1.8–2.1 × 1.2–1.4 mm, smooth to rugose. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
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Cyathia | peduncle 0.9–1.5 mm. |
solitary at distal nodes; peduncle 0.9–3 mm. |
2n | = 28. |
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Euphorbia heterophylla |
Euphorbia cordifolia |
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Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting summer–fall. |
Habitat | Disturbed areas, roadsides. | Xeric oak-pine scrub, pine-barrens, sand barrens, sandy stream banks. |
Elevation | 0–500 m. (0–1600 ft.) | 0–200 m. (0–700 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
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Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia cordifolia is easily identified by its cordate to rounded leaf base and distinctive filiform stipules. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 265. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | Chamaesyce cordifolia |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Elliott: Sketch. Bot. S. Carolina 2: 656. (1824) |
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