Euphorbia heterophylla |
Euphorbia commutata |
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Mexican fireplant, painted euphorbia |
tinted euphorbia, tinted woodland spurge |
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Habit | Herbs, annual, with taproot. | Herbs, usually biennial, occasionally annual, with taproot. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
erect or ascending, decumbent and often branched near base, 10–40 cm, glabrous. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
petiole usually 5–10 mm, 0–1 mm distally; blade usually oblanceolate to obovate, rarely ovate, 5–30 × 3–10 mm, base broadly attenuate, margins entire, apex usually obtuse to rounded, occasionally slightly retuse, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
campanulate, 1.7–2.5 × 1.5–2.5 mm, glabrous; glands 4, crescent-shaped; 0.6–1 × 0.8–1.5 mm; horns divergent, 0.5–1.1 mm. |
Staminate flowers | 8–15. |
9–15. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.9–1.3 mm, 2-fid. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
ovoid-globose, 2.5–3.2 × 3 mm, slightly lobed; cocci rounded to ± flattened, smooth, glabrous; columella 1.5–2 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
white to gray, broadly oblong-elliptic to ovoid or nearly globose, 1.5–2 × 1.3–1.6 mm, strongly small-pitted; caruncle irregularly winglike, conic, 0.6–1 × 0.7–0.9 mm. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
arrangement: terminal pleiochasial branches (2–)3(–4), 1–3+ times 2-branched; pleiochasial bracts similar in shape and size to distal leaves; dichasial bracts distinct to basally subconnate, not imbricate, widely ovate, rhombic, or reniform, base cordate, rarely slightly perfoliate, margins entire, apex obtuse to rounded, mucronulate; axillary cymose branches 0–5. |
Cyathia | peduncle 0.9–1.5 mm. |
peduncle 0.5–1 mm. |
2n | = 28. |
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Euphorbia heterophylla |
Euphorbia commutata |
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Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting spring–summer. |
Habitat | Disturbed areas, roadsides. | Bottomland and upland forests, bluffs and ledges, stream banks, glades, rarely fen margins. |
Elevation | 0–500 m. (0–1600 ft.) | 50–1000 m. (200–3300 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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AL; AR; FL; GA; IA; IL; IN; KY; LA; MD; MI; MO; MS; NC; OH; OK; PA; SC; TN; VA; WI; WV; ON
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Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 300. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | Galarhoeus austrinus, G. commutatus, Tithymalus commutatus |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Engelmann: in A. Gray, Manual ed. 2, 389. (1856) |
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