Euphorbia heterophylla |
Euphorbia cinerascens |
|
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Mexican fireplant, painted euphorbia |
ashy sandmat |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
prostrate to decumbent, mat-forming, 5–30 cm, appressed wooly, strigillose, or short-sericeous. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
opposite; stipules distinct, subulate, 0.2–0.5 mm, appressed wooly to sericeous; petiole 0.3–0.8 mm, appressed wooly to sericeous; blade ovate to elliptic, 1.5–5.5 × 1.3–4 mm, base asymmetric, obtuse to hemicordate, margins entire often reddish, apex usually obtuse, occasionally acute (young leaves), surfaces sericeous to strigillose or slightly pilose, adaxial surface often glabrous; 3-veined from base but only midvein conspicuous. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
turbinate, 0.8–1.3 × 1.2–2 mm, appressed wooly, sericeous, or strigillose; glands 4, purple-black, elliptic to oblong, 0.2–0.3 × 0.4–0.6 mm; appendages absent or reddish pink, forming narrow rim around distal margin of gland, 0–0.1 × 0–0.6 mm, distal margin entire, crenulate, or erose. |
Staminate flowers | 8–15. |
15–20. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary canescent; styles 0.3–0.5 mm, 2-fid at apex. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
subglobose to broadly ovoid, 1.3–1.7 × 1.5–1.8 mm, canescent; columella 1–1.3 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
white to pinkish or light brown, ovoid, 4-angled in cross section, 1–1.4 × 0.6–0.8 mm, smooth to rugulose or rarely with 1–2 inconspicuous transverse ridges. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
|
Cyathia | peduncle 0.9–1.5 mm. |
solitary at distal nodes; peduncle 0.2–0.3 mm. |
2n | = 28. |
= 32. |
Euphorbia heterophylla |
Euphorbia cinerascens |
|
Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting year-round (mostly spring–fall). |
Habitat | Disturbed areas, roadsides. | Desert scrub, oak and juniper woodlands, thorn scrub, shrublands, grasslands, frequently on limestone substrates. |
Elevation | 0–500 m. (0–1600 ft.) | 70–1400 m. (200–4600 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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TX; Mexico |
Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the flora area, Euphorbia cinerascens is found only in southern and western Texas. In Mexico, it is found from Chihuahua east to Tamaulipas, south to Guanajuato. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 264. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | Chamaesyce cinerascens |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) |
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