Euphorbia heterophylla |
Euphorbia bifurcata |
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Mexican fireplant, painted euphorbia |
fork spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, annual, with slender, fibrous taproot. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
erect, 20–70 cm, glabrous or with few scattered spreading hairs; branches arcuate, branching appearing dichotomous. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
usually alternate, occasionally opposite at proximalmost node; petiole 15–49 mm, glabrous; blade usually ovate, rarely oblong or elliptic, 13–54 × 7–38 mm, base usually rounded to broadly cuneate, rarely truncate, margins finely serrulate, apex obtuse, surfaces glabrous or with few scattered hairs; venation pinnate, midvein prominent. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
tubular or obconic, 1–1.7 × 0.7–1.4 mm, glabrous except for few hairs on lobes; involucral lobes divided into several linear, smooth lobes; glands 1(–3), greenish, sessile and broadly attached, 0.3–0.4 × 0.4–0.8 mm, opening oblong to subcircular, glabrous; appendages petaloid, white, elliptic, oblong, transversely oblong, or forming thin, lunate rim on gland margin, not incurved and covering glands, 0.3–0.9 × 0.6–1.3 mm, entire, undulate or slightly lobed, glabrous. |
Staminate flowers | 8–15. |
20–30. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.6–1 mm, 2-fid 1/2 length. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
oblate, 2.8–3.1 × 3.6–4.5 mm, glabrous; columella 1.9–2.4 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
brown to blackish, ovoid, rounded in cross section, 1.9–2.4 × 1.5–1.8 mm, irregularly and coarsely tuberculate; caruncle absent or rudimentary. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
arrangement: terminal dichasial branches 2, few-branched (weakly defined); pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves; dichasial bracts smaller than distal leaves, often white at base. |
Cyathia | peduncle 0.9–1.5 mm. |
peduncle 0.9–3.5(–6.2) mm. |
2n | = 28. |
|
Euphorbia heterophylla |
Euphorbia bifurcata |
|
Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting summer–fall. |
Habitat | Disturbed areas, roadsides. | Riparian areas with cottonwoods and willows, pinyon pine woodlands, pine-oak forests, Douglas fir forests with pines. |
Elevation | 0–500 m. (0–1600 ft.) | 1900–2300 m. (6200–7500 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
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NM; TX; Mexico; Central America (Guatemala) |
Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia bifurcata is found in the mountains of southern New Mexico (Doña Ana, Grant, Lincoln, Otero, and Sierra counties) and trans-Pecos Texas (Brewster, Jeff Davis, and Presidio counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 319. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859) |
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