Euphorbia heterophylla |
Euphorbia angusta |
|
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Mexican fireplant, painted euphorbia |
Blackfoot sandmat, narrow-leaf spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | erect-ascending, 20–100 cm, sparsely pilose to villous; branches ± straight. |
erect, 12–43 cm, uniformly strigose. |
Leaves | usually alternate, occasionally opposite proximally; petiole 10–50 mm, pilose; blade narrowly lanceolate to elliptic or broadly obovate (then usually pandurate and 4-lobed), often polymorphic on single plants, 30–200 × 20–140 mm, base acute, margins sparsely glandular-serrulate, hirtellous, flat, apex acute to obtuse, abaxial surface sparsely appressed-pilose, adaxial surface sparsely pilosulous to glabrate; venation pinnate, midvein prominent. |
opposite; stipules distinct, linear-subulate or nodiform to papilliform nodiform to papilliform stipules often reddish brown, 0.1–0.7 mm, strigose; petiole 0.3–1.2 mm, strigose; blade: proximal ovate to ovate-elliptic, distal linear to elliptic-linear, 7–41 × 2–5 mm, distal leaf blades more than 6 times as long as wide, base asymmetric, cuneate to rounded, margins entire, often involute on drying, apex acute, surfaces usually short strigose, occasionally glabrous adaxially; venation pinnate, only midvein conspicuous. |
Involucre | usually campanulate, occasionally nearly hemispheric, 1.5–1.9 × 1.2–1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.4 × 1–1.2 mm, opening circular (occasionally flattened from pressing), with annular rim, glabrous; appendages absent. |
turbinate to campanulate-turbinate, 1–1.5 × 1–1.4 mm, strigose; glands 4, green to yellow-green, concave, narrowly oblong, 0.2–0.4 × 0.4–0.7 mm; appendages white, flabellate, 0.5–1.1 × 0.3–0.5 mm, distal margin shallowly and irregularly toothed. |
Staminate flowers | 8–15. |
16–26. |
Pistillate flowers | ovary glabrous or puberulent; styles 0.8–1.3 mm, 2-fid 1/2 to nearly entire length. |
ovary strigose; styles 0.3–0.5 mm, 2-fid at apex to almost 1/2 length. |
Capsules | broadly ovoid, 2.8–3.8 × 4–5.3 mm, 3-lobed, usually glabrous, rarely sparsely puberulent; columella 2.1–2.8 mm. |
broadly ovoid, 2–2.6 × 2.5–3 mm, strigose; columella 1.6–2.2 mm. |
Seeds | brown-gray to ashy gray, broadly deltoid, 2.4–2.8 × 1.9–2.4 mm, angular in cross section, dorsal face strongly acute-carinate, tuberculate, with broad rounded tubercles in 2 rows; caruncle 0.1 mm. |
white, ovoid, 4-angled in cross section, 1.7–2.2 × 1.1–1.2 mm, transversely low-ridged or wrinkled. |
Cyathial | arrangement: terminal dichasial branches usually 2, occasionally reduced to congested cyme, 1–2-branched (often congested and difficult to discern); pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves; dichasial bracts highly reduced, rarely absent in highly congested clusters. |
|
Cyathia | peduncle 0.9–1.5 mm. |
solitary at distal nodes; peduncle 1.1–2.4 mm. |
2n | = 28. |
= 28. |
Euphorbia heterophylla |
Euphorbia angusta |
|
Phenology | Flowering and fruiting nearly year-round. | Flowering and fruiting early spring–fall. |
Habitat | Disturbed areas, roadsides. | Rocky limestone soils. |
Elevation | 0–500 m. (0–1600 ft.) | 400–1200 m. (1300–3900 ft.) |
Distribution |
Mexico; Central America; South America [Introduced, Ala., Ariz., Calif., Fla., Ga., La., Miss., N.Mex., Tex.; introduced also in Eurasia, Africa]
|
TX; Mexico (Coahuila) |
Discussion | Euphorbia heterophylla occurs from the southern United States, where it is likely naturalized, south through Mexico and Central America to South America. Because of its weediness, the precise native range in tropical and subtropical parts of the New World is not well understood. It has become widely established also in warm areas of the Old World. Leaf shape in this species is highly polymorphic within both populations and individuals. Euphorbia heterophylla can appear superficially similar to E. cyathophora but differs in its stipitate, circular involucral glands and its floral bracts that are at most very pale at the base (never colored as is typical in E. cyathophora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia angusta, which in the flora area is known from the trans-Pecos region to the Edwards Plateau, is easily recognized by its erect habit, linear leaves, and relatively showy involucral gland appendages with toothed margins. The species is closely related to E. acuta and the Mexican endemic E. johnstonii Mayfield (M. H. Mayfield 1991); it is not only morphologically distinctive but is also the only species in sect. Anisophyllum with C3 photosynthesis (R. F. Sage et al. 2011; T. L. Sage et al. 2011; G. L. Webster 1975). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 323. | FNA vol. 12, p. 260. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. geniculata, Poinsettia geniculata, P. heterophylla | Chamaesyce angusta |
Name authority | Linnaeus: Sp. Pl. 1: 453. (1753) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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