Flora of North America species comparison

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Euphorbia helioscopia

Euphorbia yaquiana

euphorbe réveille-matin, mad woman's milk, summer spurge, sun spurge, wart spurge, wartweed

hairy Mojave spurge
Habit Herbs, annual, with taproot. Herbs, perennial, with thick rootstock.
Stems

erect, unbranched or branched, 5–45 cm, usually glabrous or sparsely pilose.

slender, erect or ascending, sometimes sinuous, densely branched near base, 10–50 cm, moderately to densely puberulent to lanulose.
Leaves

petiole absent or to 0.5 mm;

blade obovate-spatulate, 4–40 × 2–25 mm, base cuneate, attenuate, or auriculate, margins serrulate, apex rounded, surfaces glabrous;

venation pinnate, midvein prominent.

petiole 0–1 mm;

blade usually lanceolate or elliptic-lanceolate, sometimes slightly oblanceolate, 8–30 × 6–14 mm, base usually acute, occasionally short-attenuate, rarely obtuse, margins entire, apex usually acute, occasionally obtuse, acuminate, or cuspidate, surfaces sparsely to moderately puberulent to lanulose;

venation pinnate, sometimes obscure, midvein prominent.
Involucre

cupulate, 1.5–2 × 0.7–1.1 mm, glabrous;

glands 4, elliptic, 0.2–0.5 × 0.5–1 mm;

horns absent.

campanulate to broadly turbinate, 2.2–3 × 2–2.5 mm, puberulent to lanulose;

glands 4, semicircular, trapezoidal, or elliptic-truncate, 0.8–1.5 × 1–2.2 mm, margins strongly crenate or dentate;

horns usually absent, if present then straight, 0.1–0.2 mm, generally equaling teeth on gland margin.
Staminate flowers

10–15.

12–20.
Pistillate flowers

ovary glabrous;

styles 0.7–1 mm, 2-fid.

ovary usually puberulent, occasionally lanulose;

styles 1–1.2 mm, 2-fid.
Capsules

depressed-globose, 2.5–4 × 3.2–4.2 mm, clearly 3-lobed;

cocci rounded, smooth, glabrous;

columella 0.9–1.1 mm.

oblong-ovoid, 3.5–4 × 3–4 mm, 3-lobed;

cocci rounded, smooth, usually puberulent, occasionally lanulose;

columella 2.5–3 mm.
Seeds

dark brown to blackish, subovoid, 1.6–2.2 × 1.5–1.9 mm, reticulate;

caruncle elliptic, 0.9–1.1 × 0.4–0.5 mm.

gray to whitish, oblong cylindric, 2–3 × 1.5–1.8 mm, irregularly shallowly pitted to almost smooth;

caruncle conic, 0.6 × 0.6 mm.
Cyathial

arrangement: terminal pleiochasial branches (3–)5, each 1–2 times 2-branched;

pleiochasial bracts obovate, wider than distal leaves;

dichasial bracts distinct, obovate or rhombic, ± oblique, base rounded, truncate, or attenuate, margins serrulate, apex rounded;

axillary cymose branches 0.

arrangement: terminal pleiochasial branches 3–5, each 1–2 times 2-branched;

pleiochasial bracts broadly ovate to subcordate, usually similar in size to, occasionally wider than, distal leaves;

dichasial bracts distinct, broadly ovate to almost reniform, base obtuse, margins entire, apex obtuse, acuminate to cuspidate;

axillary cymose branches 0–5.
Cyathia

peduncle 0.2–1 mm.

peduncle 0.3–0.8 mm.

Euphorbia helioscopia

Euphorbia yaquiana
Phenology Flowering and fruiting spring–fall. Flowering and fruiting spring–summer.
Habitat Roadsides, waste places. Ponderosa pine forests, oak-pine mixed forests, dry stream banks and beds, open scrub areas, roadsides.
Elevation 0–1400 m. (0–4600 ft.) 1000–2200 m. (3300–7200 ft.)
Distribution
from FNA
CA; CT; DC; DE; GA; ID; IL; LA; MA; MD; ME; MI; MT; NC; NH; NJ; NY; OH; OR; PA; SC; TN; TX; VA; VT; WA; WI; WY; AB; BC; NB; NF; NS; ON; PE; QC; SK; Europe; Asia; n Africa [Introduced in North America; introduced also in South America (Argentina, Chile)]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Euphorbia helioscopia was collected once in Minnesota in the late 1800s but apparently did not become established there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia yaquiana is endemic to Pima and Graham counties in southern Arizona and is known only from the Santa Catalina and Pinaleño mountains. Records of E. yaquiana from southwestern Colorado (as E. incisa var. mollis) likely represent misidentifications of E. brachycera; therefore, those disjunct occurrences have been excluded here from the distribution of E. yaquiana. Euphorbia yaquiana has often been treated as a synonym of E. schizoloba var. mollis, but molecular phylogenetic data show that it is more closely related to E. brachycera and E. chamaesula (J. A. Peirson et al. 2014).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 12, p. 303. FNA vol. 12, p. 313.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii
Synonyms Galarhoeus helioscopius, Tithymalus helioscopius E. schizoloba var. mollis, E. incisa var. mollis
Name authority Linnaeus: Sp. Pl. 1: 459. (1753) Tidestrom: Proc. Biol. Soc. Wash. 48: 41. (1935)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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