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euphorbe réveille-matin, mad woman's milk, summer spurge, sun spurge, wart spurge, wartweed

summer spurge

Habit Herbs, annual, with taproot. Herbs, perennial, with spreading rootstock.
Stems

erect, unbranched or branched, 5–45 cm, usually glabrous or sparsely pilose.

erect or ascending, unbranched, solitary or few, previous year's dead stems not persistent, 45–70 cm, usually densely puberulent to sericeous, rarely glabrous.

Leaves

petiole absent or to 0.5 mm;

blade obovate-spatulate, 4–40 × 2–25 mm, base cuneate, attenuate, or auriculate, margins serrulate, apex rounded, surfaces glabrous;

venation pinnate, midvein prominent.

alternate;

stipules to 0.1 mm;

petiole (0–)1–2 mm (or absent), densely puberulent;

blade usually linear, rarely ovate, 25–55 × 1.5–4 mm, base cuneate, margins entire, revolute, apex rounded, abaxial surface glabrous or puberulent to sericeous, adaxial surface glabrous;

venation often obscure on smaller leaves, midvein conspicuous.

Involucre

cupulate, 1.5–2 × 0.7–1.1 mm, glabrous;

glands 4, elliptic, 0.2–0.5 × 0.5–1 mm;

horns absent.

campanulate, 1.2–1.4 × 1.2–2 mm, sparsely to densely puberulent;

glands 5, green, reniform, 0.2–0.3 × 0.5–0.6 mm;

appendages white, orbiculate to oblong, (0.5–)1–1.7 × 1–1.5 mm, entire.

Staminate flowers

10–15.

20–25.

Pistillate flowers

ovary glabrous;

styles 0.7–1 mm, 2-fid.

ovary glabrous or sparsely strigose;

styles 0.5–1.1 mm, 2-fid at apex to 1/2 length.

Capsules

depressed-globose, 2.5–4 × 3.2–4.2 mm, clearly 3-lobed;

cocci rounded, smooth, glabrous;

columella 0.9–1.1 mm.

globose, 1.8–3 × 2.5–4.8 mm, glabrous or sparsely strigose;

columella 2.3–2.5 mm.

Seeds

dark brown to blackish, subovoid, 1.6–2.2 × 1.5–1.9 mm, reticulate;

caruncle elliptic, 0.9–1.1 × 0.4–0.5 mm.

light gray, ovoid, 2 × 1.2–1.3 mm, smooth or with few, very shallow depressions;

caruncle absent.

Cyathial

arrangement: terminal pleiochasial branches (3–)5, each 1–2 times 2-branched;

pleiochasial bracts obovate, wider than distal leaves;

dichasial bracts distinct, obovate or rhombic, ± oblique, base rounded, truncate, or attenuate, margins serrulate, apex rounded;

axillary cymose branches 0.

Cyathia

peduncle 0.2–1 mm.

in terminal pleiochasia;

peduncle 5–15 mm, filiform, glabrous or very sparsely puberulent to sericeous.

Euphorbia helioscopia

Euphorbia discoidalis

Phenology Flowering and fruiting spring–fall. Flowering and fruiting late spring–fall.
Habitat Roadsides, waste places. Sand hills, pine savannas, woodland borders, open fields with sandy soils.
Elevation 0–1400 m. (0–4600 ft.) 0–150 m. (0–500 ft.)
Distribution
from FNA
CA; CT; DC; DE; GA; ID; IL; LA; MA; MD; ME; MI; MT; NC; NH; NJ; NY; OH; OR; PA; SC; TN; TX; VA; VT; WA; WI; WY; AB; BC; NB; NF; NS; ON; PE; QC; SK; Europe; Asia; n Africa [Introduced in North America; introduced also in South America (Argentina, Chile)]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia helioscopia was collected once in Minnesota in the late 1800s but apparently did not become established there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

M. J. Huft (1979) remarked that Euphorbia discoidalis is uncommon west of Alabama and referred many narrow-leaved specimens from Louisiana and Texas to E. corollata. K. R. Park (1998) included them in an expanded E. discoidalis, and that is followed here. The western populations can be distinguished from E. corollata by their shorter involucral gland appendages and revolute leaf margins. Further study of these western populations is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 303. FNA vol. 12, p. 245.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Alectoroctonum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Galarhoeus helioscopius, Tithymalus helioscopius Tithymalopsis discoidalis
Name authority Linnaeus: Sp. Pl. 1: 459. (1753) Chapman: Fl. South. U.S., 401. (1860)
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