FNA: Euphorbia helioscopia vs. Euphorbia dendroides

	Euphorbia helioscopia	Euphorbia dendroides
	euphorbe réveille-matin, mad woman's milk, summer spurge, sun spurge, wart spurge, wartweed	tree spurge
Habit	Herbs, annual, with taproot.	Shrubs, dendroid, with large rootstock.
Stems	erect, unbranched or branched, 5–45 cm, usually glabrous or sparsely pilose.	erect, stout, densely branched, to 200 cm, glabrous, bark usually reddish and glossy toward tip of branches.
Leaves	petiole absent or to 0.5 mm; blade obovate-spatulate, 4–40 × 2–25 mm, base cuneate, attenuate, or auriculate, margins serrulate, apex rounded, surfaces glabrous; venation pinnate, midvein prominent.	petiole absent; blade linear-lanceolate to oblong-lanceolate, 20–65 × 3–8 mm, base attenuate, margins entire, apex obtuse to acute, sometimes mucronulate, surfaces glabrous; venation pinnate, midvein prominent, base attenuate.
Involucre	cupulate, 1.5–2 × 0.7–1.1 mm, glabrous; glands 4, elliptic, 0.2–0.5 × 0.5–1 mm; horns absent.	broadly turbinate to hemispheric, 3–4 × 3.5–3.8 mm, glabrous; glands 4, suborbiculate to subtrapezoidal, 1–1.5 × 1.2–1.8 mm, sometimes irregularly lobed; horns absent.
Staminate flowers	10–15.	20–25.
Pistillate flowers	ovary glabrous; styles 0.7–1 mm, 2-fid.	ovary glabrous; styles 2.4–3 mm, 2-fid.
Capsules	depressed-globose, 2.5–4 × 3.2–4.2 mm, clearly 3-lobed; cocci rounded, smooth, glabrous; columella 0.9–1.1 mm.	subglobose, 4–5.5 × 4–6.5 mm, deeply lobed; cocci laterally compressed, smooth, glabrous; columella 3–4 mm.
Seeds	dark brown to blackish, subovoid, 1.6–2.2 × 1.5–1.9 mm, reticulate; caruncle elliptic, 0.9–1.1 × 0.4–0.5 mm.	grayish or blackish, ovoid and strongly laterally compressed, 3–3.5 × 1.5–2 mm, smooth; caruncle semirounded and laterally compressed, 1.2–1.4 × 0.8–1.8 mm.
Cyathial	arrangement: terminal pleiochasial branches (3–)5, each 1–2 times 2-branched; pleiochasial bracts obovate, wider than distal leaves; dichasial bracts distinct, obovate or rhombic, ± oblique, base rounded, truncate, or attenuate, margins serrulate, apex rounded; axillary cymose branches 0.	arrangement: terminal pleiochasial branches 4–8, unbranched or 1–2-times 2-branched; pleiochasial bracts similar in shape to but usually shorter and wider than distal leaves; dichasial bracts distinct, yellowish, broadly ovate, orbiculate, rhombic, or reniform, base obtuse, margins entire, apex obtuse, sometimes mucronulate; axillary pleiochasial absent.
Cyathia	peduncle 0.2–1 mm.	peduncle absent.

	Euphorbia helioscopia	Euphorbia dendroides
Phenology	Flowering and fruiting spring–fall.	Flowering and fruiting fall–spring.
Habitat	Roadsides, waste places.	Coastal plains and basins, hillsides.
Elevation	0–1400 m. (0–4600 ft.)	0–500 m. (0–1600 ft.)
Distribution	CA; CT; DC; DE; GA; ID; IL; LA; MA; MD; ME; MI; MT; NC; NH; NJ; NY; OH; OR; PA; SC; TN; TX; VA; VT; WA; WI; WY; AB; BC; NB; NF; NS; ON; PE; QC; SK; Europe; Asia; n Africa [Introduced in North America; introduced also in South America (Argentina, Chile)] [WildflowerSearch map] [BONAP county map]	CA; Europe; n Africa [Introduced in North America] [BONAP county map]
Discussion	Euphorbia helioscopia was collected once in Minnesota in the late 1800s but apparently did not become established there. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Euphorbia dendroides is native to the Mediterranean region. In the flora area, the species is known from Los Angeles, Santa Barbara, and Ventura counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 12, p. 303.	FNA vol. 12, p. 302.
Parent taxa	Euphorbiaceae > Euphorbia > subg. Esula	Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms	Galarhoeus helioscopius, Tithymalus helioscopius	Tithymalus dendroides
Name authority	Linnaeus: Sp. Pl. 1: 459. (1753)	Linnaeus: Sp. Pl. 1: 462. (1753)
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, IL Wildflowers, LA Plants, MD Biodiversity, MI Flora, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Euphorbia dendroides

euphorbe réveille-matin, mad woman's milk, summer spurge, sun spurge, wart spurge, wartweed

tree spurge

Habit

Herbs, annual, with taproot.

Shrubs, dendroid, with large rootstock.

Stems

erect, unbranched or branched, 5–45 cm, usually glabrous or sparsely pilose.

erect, stout, densely branched, to 200 cm, glabrous, bark usually reddish and glossy toward tip of branches.

Leaves

petiole absent or to 0.5 mm;

blade obovate-spatulate, 4–40 × 2–25 mm, base cuneate, attenuate, or auriculate, margins serrulate, apex rounded, surfaces glabrous;

venation pinnate, midvein prominent.

petiole absent;

blade linear-lanceolate to oblong-lanceolate, 20–65 × 3–8 mm, base attenuate, margins entire, apex obtuse to acute, sometimes mucronulate, surfaces glabrous;

venation pinnate, midvein prominent, base attenuate.

Involucre

cupulate, 1.5–2 × 0.7–1.1 mm, glabrous;

glands 4, elliptic, 0.2–0.5 × 0.5–1 mm;

horns absent.

broadly turbinate to hemispheric, 3–4 × 3.5–3.8 mm, glabrous;

glands 4, suborbiculate to subtrapezoidal, 1–1.5 × 1.2–1.8 mm, sometimes irregularly lobed;

horns absent.

Staminate flowers

10–15.

20–25.

Pistillate flowers

ovary glabrous;

styles 0.7–1 mm, 2-fid.

ovary glabrous;

styles 2.4–3 mm, 2-fid.

Capsules

depressed-globose, 2.5–4 × 3.2–4.2 mm, clearly 3-lobed;

cocci rounded, smooth, glabrous;

columella 0.9–1.1 mm.

subglobose, 4–5.5 × 4–6.5 mm, deeply lobed;

cocci laterally compressed, smooth, glabrous;

columella 3–4 mm.

Seeds

dark brown to blackish, subovoid, 1.6–2.2 × 1.5–1.9 mm, reticulate;

caruncle elliptic, 0.9–1.1 × 0.4–0.5 mm.

grayish or blackish, ovoid and strongly laterally compressed, 3–3.5 × 1.5–2 mm, smooth;

caruncle semirounded and laterally compressed, 1.2–1.4 × 0.8–1.8 mm.

Cyathial

arrangement: terminal pleiochasial branches (3–)5, each 1–2 times 2-branched;

pleiochasial bracts obovate, wider than distal leaves;

dichasial bracts distinct, obovate or rhombic, ± oblique, base rounded, truncate, or attenuate, margins serrulate, apex rounded;

axillary cymose branches 0.

arrangement: terminal pleiochasial branches 4–8, unbranched or 1–2-times 2-branched;

pleiochasial bracts similar in shape to but usually shorter and wider than distal leaves;

dichasial bracts distinct, yellowish, broadly ovate, orbiculate, rhombic, or reniform, base obtuse, margins entire, apex obtuse, sometimes mucronulate;

axillary pleiochasial absent.

Cyathia

peduncle 0.2–1 mm.

peduncle absent.

Euphorbia dendroides

Phenology

Flowering and fruiting spring–fall.

Flowering and fruiting fall–spring.

Habitat

Roadsides, waste places.

Coastal plains and basins, hillsides.

Elevation

0–1400 m. (0–4600 ft.)

0–500 m. (0–1600 ft.)

Distribution

CA; CT; DC; DE; GA; ID; IL; LA; MA; MD; ME; MI; MT; NC; NH; NJ; NY; OH; OR; PA; SC; TN; TX; VA; VT; WA; WI; WY; AB; BC; NB; NF; NS; ON; PE; QC; SK; Europe; Asia; n Africa [Introduced in North America; introduced also in South America (Argentina, Chile)]

[WildflowerSearch map]

[BONAP county map]

CA; Europe; n Africa [Introduced in North America]

[BONAP county map]

Discussion

Euphorbia helioscopia was collected once in Minnesota in the late 1800s but apparently did not become established there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia dendroides is native to the Mediterranean region. In the flora area, the species is known from Los Angeles, Santa Barbara, and Ventura counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 12, p. 303.

FNA vol. 12, p. 302.

Parent taxa

Euphorbiaceae > Euphorbia > subg. Esula

Sibling taxa

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

Synonyms

Galarhoeus helioscopius, Tithymalus helioscopius

Tithymalus dendroides

Name authority

Linnaeus: Sp. Pl. 1: 459. (1753)

Linnaeus: Sp. Pl. 1: 462. (1753)

Web links

Local floras: BC, CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, IL Wildflowers, LA Plants, MD Biodiversity, MI Flora, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Euphorbia helioscopia

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