FNA: Euphorbia graminea vs. Euphorbia misera

	Euphorbia graminea	Euphorbia misera
	grassleaf spurge	cliff spurge
Habit	Herbs, usually annual, rarely perennial, with slender, rarely tuberous, taproot.	Shrubs, soft wooded, with woody rootstock.
Stems	erect or ascending, branched, 30–80(–110) cm, strigillose or glabrescent, sharply angled.	erect to ascending, often gnarled and scraggly, branched, with conspicuous knobby short shoots, 70–150 cm, puberulent-tomentose, bark grayish red to light gray.
Leaves	usually alternate, sometimes some opposite; stipules usually 0.2–0.5 mm, rarely rudimentary; petiole 0.4–5.9 mm, strigillose; blade ovate, elliptic, linear-elliptic, or oblong, 10–83 × 3–39 mm, base attenuate, rounded, or cuneate, margins entire, apex acute or obtuse, surfaces strigillose; venation occasionally obscure on narrow leaves, midvein conspicuous.	alternate, well spaced on long shoots or fasciculate on short shoots; stipules 0.6–1.1 mm; petiole 4–12(–19) mm, slender, puberulent to shortly pilose; blade oblong, ovate, orbiculate, elliptic, or obovate, 6–24 × 5–21 mm, base rounded to cuneate, margins entire, apex acute to obtuse, surfaces puberulent-tomentose; venation conspicuous.
Involucre	campanulate or obconic, 1–1.8 × 0.8–1.7 mm, glabrous or strigillose toward rim; glands (1–)2–4, yellow to greenish, elliptic or oblong, 0.1–0.3 × 0.2–0.4 mm; appendages white to tinged purple, ovate and often hoodlike or forming narrow rim around distal margin of gland, 0.3–1.6 × 0.4–0.9 mm, entire.	campanulate, 1.4–3.8 × 2.1–4.4 mm, puberulent-tomentose; glands 5, yellow to reddish, oblong to reniform, 0.7–1.3 × 1.1–2.6 mm; appendages green-yellow to yellowish or whitish, oblong to transversely oblong, 0.6–1.9 × 1.3–3.8 mm, crenulate to erose.
Staminate flowers	30–40.	40–50.
Pistillate flowers	ovary glabrous; styles 0.7–1 mm, 2-fid from 1/2 to nearly entire length.	ovary glabrous or puberulent; styles 1.6–2.7 mm, 2-fid at apex.
Capsules	ovoid-oblate, 2.5–3 × 3–3.5 mm, glabrous; columella 1.6–1.9 mm.	oblate, 4.6–5.1 × 6.1–6.7 mm, usually glabrous or glabrescent, occasionally puberulent; columella 2.8–3.6 mm.
Seeds	gray, brown, or nearly black, ovoid, circular or weakly angled in cross section, 1.5–1.7 × 1.3–1.5 mm, coarsely tuberculate with longitudinal rows of shallow pits; caruncle absent or punctiform, 0.1–0.2 mm.	grayish, subglobose to ovoid, rounded in cross section, 2.7–3.3 × 2.5–2.8 mm, foveolate; caruncle absent.
Cyathia	in usually terminal, rarely axillary, dichasia, distal dichasial bracts often white; peduncle 0.4–4.5 mm (to 15 mm at first node of inflorescence), glabrous.	usually solitary on short shoots, peduncle 1.8–10.5 mm, puberulent-tomentose.

	Euphorbia graminea	Euphorbia misera
Phenology	Flowering and fruiting year-round.	Flowering and fruiting year-round (but most prolific after winter rains).
Habitat	Disturbed, weedy, or urban areas.	Rocky soils, sometimes in crevices of vertical cliff faces, coastal scrub, maritime desert scrub, arid desert scrub.
Elevation	0–500 m. (0–1600 ft.)	0–400 m. (0–1300 ft.)
Distribution	AR; CA; FL; LA; TX; Mexico; Central America; South America [Introduced in North America; introduced also in West Indies, Asia, Pacific Islands] [WildflowerSearch map] [BONAP county map]	CA; Mexico (Baja California, Baja California Sur, Sonora) [WildflowerSearch map] [BONAP county map]
Discussion	Euphorbia graminea occurs natively from northern South America to northern Mexico. The species is a variable and taxonomically complex entity whose boundaries are not well defined and are in need of further study. Euphorbia graminea is often weedy and has recently become established in warmer areas of the southern United States, where it will likely become more common in the future. In recent years, a cultivar of E. graminea has found considerable horticultural success and is marketed under the trade name "Diamond Frost." (Discussion copyrighted by Flora of North America; reprinted with permission.)	Euphorbia misera is relatively infrequent within the flora area, known primarily from coastal southern California and the Channel Islands (although a relictual inland population occurs in the Little San Bernardino Mountains). The species has been considered worthy of conservation, but appears to be under little threat, especially in Mexico where it is frequent and often locally abundant. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 12, p. 246.	FNA vol. 12, p. 249.
Parent taxa	Euphorbiaceae > Euphorbia > sect. Alectoroctonum	Euphorbiaceae > Euphorbia > sect. Alectoroctonum
Sibling taxa	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Name authority	Jacquin: Select. Stirp. Amer. Hist., 151. (1763)	Bentham: Bot. Voy. Sulphur, 51. (1844)
Web links	Local Web sites: CalFlora, CalPhotos, LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Euphorbia misera

grassleaf spurge

cliff spurge

Habit

Herbs, usually annual, rarely perennial, with slender, rarely tuberous, taproot.

Shrubs, soft wooded, with woody rootstock.

Stems

erect or ascending, branched, 30–80(–110) cm, strigillose or glabrescent, sharply angled.

erect to ascending, often gnarled and scraggly, branched, with conspicuous knobby short shoots, 70–150 cm, puberulent-tomentose, bark grayish red to light gray.

Leaves

usually alternate, sometimes some opposite;

stipules usually 0.2–0.5 mm, rarely rudimentary;

petiole 0.4–5.9 mm, strigillose;

blade ovate, elliptic, linear-elliptic, or oblong, 10–83 × 3–39 mm, base attenuate, rounded, or cuneate, margins entire, apex acute or obtuse, surfaces strigillose;

venation occasionally obscure on narrow leaves, midvein conspicuous.

alternate, well spaced on long shoots or fasciculate on short shoots;

stipules 0.6–1.1 mm;

petiole 4–12(–19) mm, slender, puberulent to shortly pilose;

blade oblong, ovate, orbiculate, elliptic, or obovate, 6–24 × 5–21 mm, base rounded to cuneate, margins entire, apex acute to obtuse, surfaces puberulent-tomentose;

venation conspicuous.

Involucre

campanulate or obconic, 1–1.8 × 0.8–1.7 mm, glabrous or strigillose toward rim;

glands (1–)2–4, yellow to greenish, elliptic or oblong, 0.1–0.3 × 0.2–0.4 mm;

appendages white to tinged purple, ovate and often hoodlike or forming narrow rim around distal margin of gland, 0.3–1.6 × 0.4–0.9 mm, entire.

campanulate, 1.4–3.8 × 2.1–4.4 mm, puberulent-tomentose;

glands 5, yellow to reddish, oblong to reniform, 0.7–1.3 × 1.1–2.6 mm;

appendages green-yellow to yellowish or whitish, oblong to transversely oblong, 0.6–1.9 × 1.3–3.8 mm, crenulate to erose.

Staminate flowers

30–40.

40–50.

Pistillate flowers

ovary glabrous;

styles 0.7–1 mm, 2-fid from 1/2 to nearly entire length.

ovary glabrous or puberulent;

styles 1.6–2.7 mm, 2-fid at apex.

Capsules

ovoid-oblate, 2.5–3 × 3–3.5 mm, glabrous;

columella 1.6–1.9 mm.

oblate, 4.6–5.1 × 6.1–6.7 mm, usually glabrous or glabrescent, occasionally puberulent;

columella 2.8–3.6 mm.

Seeds

gray, brown, or nearly black, ovoid, circular or weakly angled in cross section, 1.5–1.7 × 1.3–1.5 mm, coarsely tuberculate with longitudinal rows of shallow pits;

caruncle absent or punctiform, 0.1–0.2 mm.

grayish, subglobose to ovoid, rounded in cross section, 2.7–3.3 × 2.5–2.8 mm, foveolate;

caruncle absent.

Cyathia

in usually terminal, rarely axillary, dichasia, distal dichasial bracts often white;

peduncle 0.4–4.5 mm (to 15 mm at first node of inflorescence), glabrous.

usually solitary on short shoots, peduncle 1.8–10.5 mm, puberulent-tomentose.

Euphorbia misera

Phenology

Flowering and fruiting year-round.

Flowering and fruiting year-round (but most prolific after winter rains).

Habitat

Disturbed, weedy, or urban areas.

Rocky soils, sometimes in crevices of vertical cliff faces, coastal scrub, maritime desert scrub, arid desert scrub.

Elevation

0–500 m. (0–1600 ft.)

0–400 m. (0–1300 ft.)

Distribution

AR; CA; FL; LA; TX; Mexico; Central America; South America [Introduced in North America; introduced also in West Indies, Asia, Pacific Islands]

[WildflowerSearch map]

[BONAP county map]

CA; Mexico (Baja California, Baja California Sur, Sonora)

[WildflowerSearch map]

[BONAP county map]

Discussion

Euphorbia graminea occurs natively from northern South America to northern Mexico. The species is a variable and taxonomically complex entity whose boundaries are not well defined and are in need of further study. Euphorbia graminea is often weedy and has recently become established in warmer areas of the southern United States, where it will likely become more common in the future. In recent years, a cultivar of E. graminea has found considerable horticultural success and is marketed under the trade name "Diamond Frost."

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia misera is relatively infrequent within the flora area, known primarily from coastal southern California and the Channel Islands (although a relictual inland population occurs in the Little San Bernardino Mountains). The species has been considered worthy of conservation, but appears to be under little threat, especially in Mexico where it is frequent and often locally abundant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 12, p. 246.

FNA vol. 12, p. 249.

Parent taxa

Euphorbiaceae > Euphorbia > sect. Alectoroctonum

Sibling taxa

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

Name authority

Jacquin: Select. Stirp. Amer. Hist., 151. (1763)

Bentham: Bot. Voy. Sulphur, 51. (1844)

Web links

Local Web sites: CalFlora, CalPhotos, LA Plants
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Euphorbia graminea

Euphorbia misera

Euphorbia graminea

Euphorbia misera