Euphorbia graminea |
Euphorbia geyeri |
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grassleaf spurge |
Geyer's sandmat, Geyer's spurge |
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Habit | Herbs, usually annual, rarely perennial, with slender, rarely tuberous, taproot. | Herbs, annual, with taproot. | ||||
Stems | erect or ascending, branched, 30–80(–110) cm, strigillose or glabrescent, sharply angled. |
prostrate or slightly ascending, loosely mat-forming, 4–25(–45) cm, glabrous. |
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Leaves | usually alternate, sometimes some opposite; stipules usually 0.2–0.5 mm, rarely rudimentary; petiole 0.4–5.9 mm, strigillose; blade ovate, elliptic, linear-elliptic, or oblong, 10–83 × 3–39 mm, base attenuate, rounded, or cuneate, margins entire, apex acute or obtuse, surfaces strigillose; venation occasionally obscure on narrow leaves, midvein conspicuous. |
opposite; stipules usually distinct, occasionally connate basally on lower side, usually deeply parted into 3 or more filiform segments, 0.7–1.5 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong to oblong-obovate or oblong-elliptic, 4–12 × 2–6 mm, base slightly asymmetric, angled or rounded, with one side usually expanded into small, rounded auricle, margins entire, apex usually truncate, occasionally emarginate, abaxial surface pale grayish green, both surfaces glabrous; only midvein conspicuous or venation obscurely pinnate (larger leaves). |
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Involucre | campanulate or obconic, 1–1.8 × 0.8–1.7 mm, glabrous or strigillose toward rim; glands (1–)2–4, yellow to greenish, elliptic or oblong, 0.1–0.3 × 0.2–0.4 mm; appendages white to tinged purple, ovate and often hoodlike or forming narrow rim around distal margin of gland, 0.3–1.6 × 0.4–0.9 mm, entire. |
broadly campanulate, 1–1.5 × 0.7–0.9 mm, glabrous; glands 4, green to reddish, elliptic-oblong to nearly circular, slightly cupped to folded, 0.2–0.4 × 0.2–0.6 mm; appendages rudimentary to absent or white to reddish-tinged, usually rounded, sometimes pointed, (0–)0.5–1 × (0–)0.1–1.2 mm, distal margin entire or slightly toothed. |
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Staminate flowers | 30–40. |
5–20. |
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Pistillate flowers | ovary glabrous; styles 0.7–1 mm, 2-fid from 1/2 to nearly entire length. |
ovary glabrous; styles 0.2–0.6 mm, 2-fid nearly 1/2 length. |
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Capsules | ovoid-oblate, 2.5–3 × 3–3.5 mm, glabrous; columella 1.6–1.9 mm. |
globose-ovoid, 1.5–2 × 1.5–3 mm, glabrous; columella 1.5–1.9 mm. |
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Seeds | gray, brown, or nearly black, ovoid, circular or weakly angled in cross section, 1.5–1.7 × 1.3–1.5 mm, coarsely tuberculate with longitudinal rows of shallow pits; caruncle absent or punctiform, 0.1–0.2 mm. |
ashy white, ovoid, terete to bluntly subangled in cross section, 1.1–1.7 × 0.9–1.2 mm, smooth, with smooth brown line from top to bottom on adaxial side. |
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Cyathia | in usually terminal, rarely axillary, dichasia, distal dichasial bracts often white; peduncle 0.4–4.5 mm (to 15 mm at first node of inflorescence), glabrous. |
solitary or in small, cymose clusters at distal nodes; peduncle 1–2 mm. |
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Euphorbia graminea |
Euphorbia geyeri |
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Phenology | Flowering and fruiting year-round. | |||||
Habitat | Disturbed, weedy, or urban areas. | |||||
Elevation | 0–500 m. (0–1600 ft.) | |||||
Distribution |
AR; CA; FL; LA; TX; Mexico; Central America; South America [Introduced in North America; introduced also in West Indies, Asia, Pacific Islands]
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CO; IA; IL; IN; KS; LA; MI; MN; MO; MS; MT; ND; NE; NM; OK; SD; TX; UT; WI; WY; MB; n Mexico
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Discussion | Euphorbia graminea occurs natively from northern South America to northern Mexico. The species is a variable and taxonomically complex entity whose boundaries are not well defined and are in need of further study. Euphorbia graminea is often weedy and has recently become established in warmer areas of the southern United States, where it will likely become more common in the future. In recent years, a cultivar of E. graminea has found considerable horticultural success and is marketed under the trade name "Diamond Frost." (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). The two varieties of Euphorbia geyeri have been distinguished in large part by the presence of conspicuous involucral gland appendages in var. geyeri and the lack of appendages in var. wheeleriana. The two varieties are recognized here, but the variation in the size and presence of involucral gland appendages in the closely related E. bombensis suggests that this might be a somewhat variable character in this group of species. Euphorbia geyeri is widespread throughout the central United States in sandy soils. Populations at the eastern edge of the range are often considered adventive (for example, sandy soils along railroad grades in Michigan). Euphorbia geyeri resembles E. glyptosperma (both being entirely glabrous), but that species has serrulate leaves (near the apex) and strongly angled, transverse-ridged seeds whereas E. geyeri has entire leaves and smooth, rounded seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 246. | FNA vol. 12, p. 268. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce geyeri | |||||
Name authority | Jacquin: Select. Stirp. Amer. Hist., 151. (1763) | Engelmann: Boston J. Nat. Hist. 5: 260. (1845) | ||||
Web links |