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corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge

weak spurge

Habit Herbs, annual, with taproot. Herbs, annual, with taproot.
Stems

prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous.

erect, unbranched, 7–20 cm, glabrous.

Leaves

opposite;

stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous;

petiole 0.2–2 mm, glabrous;

blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally.

petiole 1–2 mm, reduced distally;

blade spatulate-cuneate, 8–10 × 4–5 mm (greatly reduced in size proximally), base cuneate, margins entire, apex rounded to emarginate or obcordate, surfaces glabrous;

venation pinnate.

Involucre

obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous;

glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm;

appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed.

campanulate, 0.8–1.1 × 0.7–1 mm, glabrous;

glands 4, elliptic to trapezoidal, 0.3–0.6 × 0.6–1.2 mm;

horns divergent, 0.5–1 mm.

Staminate flowers

1–5.

10–15.

Pistillate flowers

ovary glabrous;

styles 0.1–0.3 mm, 2-fid 1/2 length.

ovary smooth, glabrous;

styles 0.6–1 mm, 2-fid.

Capsules

broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous;

columella 1.3–1.5 mm.

depressed-globose, 1.8–2.2 × 2.2–2.9 mm, slightly lobed;

cocci rounded to slightly flattened, smooth, glabrous;

columella 1.5–1.8 mm.

Seeds

with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel.

reddish brown to brown, often glaucous, oblong, 1.3–1.4 × 0.8–0.9 mm, abaxial faces with 15–20 shallow pits or almost smooth, adaxial faces with 4–6 large shallow pits or irregular oblong grooves;

caruncle conic, hat-shaped, 0.3–0.4 × 0.4–0.6 mm.

Cyathia

solitary or in small, cymose clusters at distal nodes;

peduncle 0.5–1.6 mm.

peduncle 0.2–0.6 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, each 1–3(–4) times 2-branched;

pleiochasial bracts obovate, similar in size to distal leaves;

dichasial bracts distinct or basally subconnate, not imbricate, triangular-ovate, base truncate or cordate, margins entire, apex mucronate;

axillary cymose branches 1–4.

2n

= 22.

Euphorbia glyptosperma

Euphorbia tetrapora

Phenology Flowering and fruiting early summer–fall. Flowering and fruiting spring.
Habitat Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. Sandy soils, dry open woods.
Elevation 0–1800 m. (0–5900 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; OK; TX
[BONAP county map]
Discussion

Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia tetrapora is endemic to a portion of the western Gulf coastal plain. D. S. Correll and M. C. Johnston (1970) included Alabama and Georgia in the distribution of this species as well, probably due to Engelmann’s citation of a Georgia specimen from the herbarium of Samuel Boykin. Whether the Boykin specimen came from Georgia, where Boykin was based, is unclear. Because no records to support its occurrence in the eastern Gulf coastal plain (Alabama or Georgia) have been found, those states are here excluded from the distribution of E. tetrapora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 269. FNA vol. 12, p. 311.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce glyptosperma Tithymalus tetraporus
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 191. (1859)
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