Euphorbia glyptosperma |
Euphorbia ipecacuanhae |
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corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge |
American ipecac, Carolina ipecac, ipecac spurge |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with deep, stout rootstock. |
Stems | prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous. |
decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
Leaves | opposite; stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous; petiole 0.2–2 mm, glabrous; blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally. |
alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
Involucre | obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous; glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm; appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed. |
yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
Staminate flowers | 1–5. |
10–20. |
Pistillate flowers | ovary glabrous; styles 0.1–0.3 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
Capsules | broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous; columella 1.3–1.5 mm. |
globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
Seeds | with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel. |
white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
Cyathia | solitary or in small, cymose clusters at distal nodes; peduncle 0.5–1.6 mm. |
usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
2n | = 22. |
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Euphorbia glyptosperma |
Euphorbia ipecacuanhae |
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Phenology | Flowering and fruiting early summer–fall. | Flowering and fruiting early spring–early summer. |
Habitat | Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. |
Elevation | 0–1800 m. (0–5900 ft.) | 0–150 m. (0–500 ft.) |
Distribution |
AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
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CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
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Discussion | Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 269. | FNA vol. 12, p. 247. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce glyptosperma | E. arundelana, Tithymalopsis ipecacuanhae |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) | Linnaeus: Sp. Pl. 1: 455. (1753) |
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