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corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge

grassleaf spurge

Habit Herbs, annual, with taproot. Herbs, usually annual, rarely perennial, with slender, rarely tuberous, taproot.
Stems

prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous.

erect or ascending, branched, 30–80(–110) cm, strigillose or glabrescent, sharply angled.

Leaves

opposite;

stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous;

petiole 0.2–2 mm, glabrous;

blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally.

usually alternate, sometimes some opposite;

stipules usually 0.2–0.5 mm, rarely rudimentary;

petiole 0.4–5.9 mm, strigillose;

blade ovate, elliptic, linear-elliptic, or oblong, 10–83 × 3–39 mm, base attenuate, rounded, or cuneate, margins entire, apex acute or obtuse, surfaces strigillose;

venation occasionally obscure on narrow leaves, midvein conspicuous.

Involucre

obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous;

glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm;

appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed.

campanulate or obconic, 1–1.8 × 0.8–1.7 mm, glabrous or strigillose toward rim;

glands (1–)2–4, yellow to greenish, elliptic or oblong, 0.1–0.3 × 0.2–0.4 mm;

appendages white to tinged purple, ovate and often hoodlike or forming narrow rim around distal margin of gland, 0.3–1.6 × 0.4–0.9 mm, entire.

Staminate flowers

1–5.

30–40.

Pistillate flowers

ovary glabrous;

styles 0.1–0.3 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.7–1 mm, 2-fid from 1/2 to nearly entire length.

Capsules

broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous;

columella 1.3–1.5 mm.

ovoid-oblate, 2.5–3 × 3–3.5 mm, glabrous;

columella 1.6–1.9 mm.

Seeds

with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel.

gray, brown, or nearly black, ovoid, circular or weakly angled in cross section, 1.5–1.7 × 1.3–1.5 mm, coarsely tuberculate with longitudinal rows of shallow pits;

caruncle absent or punctiform, 0.1–0.2 mm.

Cyathia

solitary or in small, cymose clusters at distal nodes;

peduncle 0.5–1.6 mm.

in usually terminal, rarely axillary, dichasia, distal dichasial bracts often white;

peduncle 0.4–4.5 mm (to 15 mm at first node of inflorescence), glabrous.

2n

= 22.

Euphorbia glyptosperma

Euphorbia graminea

Phenology Flowering and fruiting early summer–fall. Flowering and fruiting year-round.
Habitat Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. Disturbed, weedy, or urban areas.
Elevation 0–1800 m. (0–5900 ft.) 0–500 m. (0–1600 ft.)
Distribution
from FNA
AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CA; FL; LA; TX; Mexico; Central America; South America [Introduced in North America; introduced also in West Indies, Asia, Pacific Islands]
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia graminea occurs natively from northern South America to northern Mexico. The species is a variable and taxonomically complex entity whose boundaries are not well defined and are in need of further study. Euphorbia graminea is often weedy and has recently become established in warmer areas of the southern United States, where it will likely become more common in the future. In recent years, a cultivar of E. graminea has found considerable horticultural success and is marketed under the trade name "Diamond Frost."

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 269. FNA vol. 12, p. 246.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Alectoroctonum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce glyptosperma
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) Jacquin: Select. Stirp. Amer. Hist., 151. (1763)
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