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corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge

Fendler's sandmat

Habit Herbs, annual, with taproot. Herbs, perennial, with woody, thickened taproot.
Stems

prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous.

usually prostrate, decumbent, or ascending, very rarely erect, often densely clustered from top of woody crown, 5–12 cm, glabrous.

Leaves

opposite;

stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous;

petiole 0.2–2 mm, glabrous;

blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally.

usually opposite, rarely whorled;

stipules distinct, narrowly linear, 0.5–1 mm, glabrous;

petiole 0.5–1 mm, glabrous;

blade usually orbiculate to ovate, rarely almost lanceolate, 3–8 × 2.5–7 mm, base slightly asymmetric, slightly cordate to rounded or obtuse, margins entire, apex rounded to obtuse, surfaces glabrous; obscurely 3–5-veined from base, only midvein conspicuous.

Involucre

obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous;

glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm;

appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed.

campanulate to turbinate or broadly cupulate, 1.1–1.7 × 1.2–1.8 mm, glabrous;

glands 4, yellow-green to reddish, elliptic to oblong, 0.2–0.5 × 0.4–0.9 mm;

appendages absent or white, rarely pink, often unequal, lunate to flabellate or sometimes forming crenate margin along gland, (0–)0.1–0.6 × (0–)0.5–1.5 mm, distal margin entire or toothed.

Staminate flowers

1–5.

25–35.

Pistillate flowers

ovary glabrous;

styles 0.1–0.3 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.3–0.4 mm, 2-fid 1/2 length.

Capsules

broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous;

columella 1.3–1.5 mm.

depressed-globose, 2–2.4 × 2.2–2.5 mm, glabrous;

columella 1.7–2.1 mm.

Seeds

with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel.

white, ovoid-pyramidal, prominently 4-angled in cross section, 1.7–2 × 1–1.2 mm, smooth to slightly wrinkled.

Cyathia

solitary or in small, cymose clusters at distal nodes;

peduncle 0.5–1.6 mm.

solitary at distal nodes;

peduncle 0.7–1.2 mm.

2n

= 22.

= 28.

Euphorbia glyptosperma

Euphorbia fendleri

Phenology Flowering and fruiting early summer–fall. Flowering and fruiting spring–fall.
Habitat Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. Mountain slopes, desert scrub, pinyon-juniper woodlands, hills, canyons, grasslands, washes, roadsides, dry crevices in limestone, often in gravel and sand.
Elevation 0–1800 m. (0–5900 ft.) 500–2600 m. (1600–8500 ft.)
Distribution
from FNA
AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; KS; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Coahuila)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia fendleri is similar to E. chaetocalyx and may sometimes be confused with that species. Its prostrate to decumbent or ascending stems and small, ovate to orbiculate leaves distinguish it from E. chaetocalyx. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. fendleri from E. chaetocalyx, but those characters appear highly variable and of little taxonomic utility; somewhat intermediate individuals occur in western Arizona, New Mexico, and Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 269. FNA vol. 12, p. 267.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce glyptosperma Chamaesyce fendleri
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 175. (1857)
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