Euphorbia glyptosperma |
Euphorbia antisyphilitica |
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corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge |
candelilla, wax plant |
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Habit | Herbs, annual, with taproot. | Shrubs, with much-branched, fleshy rootstock. |
Stems | prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous. |
erect, few branched, 25–50(–100) cm, glabrous or puberulent, pencil-like, in age covered with flaky, exfoliating layer of wax. |
Leaves | opposite; stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous; petiole 0.2–2 mm, glabrous; blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally. |
alternate usually caducous, sometimes persisting; stipules 0.4–0.5 mm; petiole absent; blade ovate to deltate-subulate, 2.5–4 × 1 mm, thick, fleshy, base usually rounded and swollen, rarely cuneate, margins entire, apex acute, surfaces puberulent, adaxial sometimes canescent; venation inconspicuous. |
Involucre | obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous; glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm; appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed. |
campanulate, 1.6–2.2 × 1.6–1.9 mm, puberulent to canescent; glands 5, pinkish, narrowly oblong to reniform, 0.3–0.4 × 0.8–1 mm; appendages white to pink, ovate, oblong, or transversely oblong, 1.3–2.5 × 1.4–2.5 mm, usually erose, rarely entire. |
Staminate flowers | 1–5. |
50–70. |
Pistillate flowers | ovary glabrous; styles 0.1–0.3 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.9–1.1 mm, 2-fid nearly entire length. |
Capsules | broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous; columella 1.3–1.5 mm. |
oblong to ovoid, 3.9–4.2 × 3.6–3.9 mm, glabrous; columella 3.1–3.3 mm. |
Seeds | with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel. |
whitish gray, narrowly ovoid, 2.4–3.1 × 1.4–1.6 mm, irregularly rugose-tuberculate; caruncle crescent-shaped, 0.3–0.6 × 0.6–0.8 mm. |
Cyathia | solitary or in small, cymose clusters at distal nodes; peduncle 0.5–1.6 mm. |
in axillary congested cymes, near branch tips or solitary at distal nodes; peduncle 0–1 mm, lanulose. |
2n | = 22. |
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Euphorbia glyptosperma |
Euphorbia antisyphilitica |
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Phenology | Flowering and fruiting early summer–fall. | Flowering and fruiting year-round in response to sufficient rainfall. |
Habitat | Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. | Desert scrub, frequently on limestone substrates. |
Elevation | 0–1800 m. (0–5900 ft.) | 100–1200 m. (300–3900 ft.) |
Distribution |
AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
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NM; TX; Mexico
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Discussion | Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia antisyphilitica is the only pencil-stemmed species of Euphorbia occurring in the flora area. The species is characteristic of the Chihuahuan Desert scrub of Mexico from Chihuahua and Coahuila south to Hidalgo and Querétaro, and barely enters into the United States in southern New Mexico (Doña Ana and Lincoln counties) and southwest (Brewster, Hudspeth, Presidio, and Terrell counties) and south (Starr and Webb counties) Texas. The stems are covered in a conspicuous coat of exfoliating wax, and the plants historically have been harvested for this product, although the practice is much less prevalent now. The specific epithet refers to its traditional medicinal use in treating sexually transmitted infections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 269. | FNA vol. 12, p. 243. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce glyptosperma | |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) | Zuccarini: Flora 15(2, Beibl.): 58. (1832) |
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