Euphorbia glyptosperma |
Euphorbia albomarginata |
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corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge |
rattlesnake sandmat, rattlesnake weed, white-margin sandmat or sandwort, whitemargin sandmat |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous. |
prostrate, occasionally mat-forming, frequently rooting at nodes, 10–80 cm, glabrous. |
Leaves | opposite; stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous; petiole 0.2–2 mm, glabrous; blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally. |
opposite; stipules connate into conspicuous, deltate or ovate scale, white, 0.4–1(–2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 3–8(–15) × 3–7 mm, base strongly asymmetric, obtuse to hemicordate, margins whitish, entire, apex obtuse, rarely mucronulate, surfaces often with red blotch in center, glabrous; 3-veined from base but usually only midvein conspicuous. |
Involucre | obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous; glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm; appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed. |
campanulate, 0.8–1.1 × 0.9–2 mm, glabrous; glands 4, greenish yellow to red, usually oblong to reniform, rarely subcircular, 0.2–0.5 × (0.2–)0.3–0.8 mm; appendages white to pink, flabellate to oblong, 0.3–1 × 0.6–1.3 mm, distal margin entire or crenulate to erose. |
Staminate flowers | 1–5. |
15–30. |
Pistillate flowers | ovary glabrous; styles 0.1–0.3 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.3–0.7 mm, 2-fid nearly entire length. |
Capsules | broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous; columella 1.3–1.5 mm. |
broadly ovoid, 1.1–2.3 × 1.2–2 mm, glabrous; columella 1.1–1.6 mm. |
Seeds | with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel. |
white to gray or brownish red, oblong-ovoid, 4-angled in cross section, 1–1.7 × 0.5–0.8 mm, smooth. |
Cyathia | solitary or in small, cymose clusters at distal nodes; peduncle 0.5–1.6 mm. |
solitary at distal nodes; peduncle 1–4 mm. |
2n | = 22. |
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Euphorbia glyptosperma |
Euphorbia albomarginata |
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Phenology | Flowering and fruiting early summer–fall. | Flowering and fruiting year-round. |
Habitat | Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. | Disturbed areas in desert scrub, grasslands, mesquite woodlands, chaparral. |
Elevation | 0–1800 m. (0–5900 ft.) | 0–2300 m. (0–7500 ft.) |
Distribution |
AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
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AZ; CA; CO; NM; NV; OK; TX; UT; Mexico [Introduced in Pacific Islands (Hawaii)]
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Discussion | Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia albomarginata is native to northern and central Mexico and the southwestern and south-central United States. The species occurs in a variety of habitats in western North America and in some areas is quite weedy. It has been recorded as a waif in Louisiana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 269. | FNA vol. 12, p. 259. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce glyptosperma | Chamaesyce albomarginata |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) | Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 174. (1857) |
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