Euphorbia geyeri |
Euphorbia chaetocalyx |
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Geyer's sandmat, Geyer's spurge |
bristlecup sandmat |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with woody, thickened taproot. | ||||||||
Stems | prostrate or slightly ascending, loosely mat-forming, 4–25(–45) cm, glabrous. |
usually erect, rarely slightly decumbent, often densely clustered from top of woody crown, 3–15 cm, glabrous. |
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Leaves | opposite; stipules usually distinct, occasionally connate basally on lower side, usually deeply parted into 3 or more filiform segments, 0.7–1.5 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong to oblong-obovate or oblong-elliptic, 4–12 × 2–6 mm, base slightly asymmetric, angled or rounded, with one side usually expanded into small, rounded auricle, margins entire, apex usually truncate, occasionally emarginate, abaxial surface pale grayish green, both surfaces glabrous; only midvein conspicuous or venation obscurely pinnate (larger leaves). |
opposite; stipules distinct, narrowly linear, usually entire, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to lanceolate or oblong- or linear-lanceolate, 3–11 × 0.8–3(–5) mm, base slightly asymmetric, short-tapered, occasionally one side slightly rounded, margins entire, apex acute or short-acuminate, surfaces glabrous; only midvein conspicuous. |
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Involucre | broadly campanulate, 1–1.5 × 0.7–0.9 mm, glabrous; glands 4, green to reddish, elliptic-oblong to nearly circular, slightly cupped to folded, 0.2–0.4 × 0.2–0.6 mm; appendages rudimentary to absent or white to reddish-tinged, usually rounded, sometimes pointed, (0–)0.5–1 × (0–)0.1–1.2 mm, distal margin entire or slightly toothed. |
campanulate to turbinate, 0.8–1.4 × 0.8–1 mm, glabrous; glands 4, yellow-brown to reddish, concave or convex, elliptic or oval, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, lanceolate-deltate to straplike, 0.2–1.1 × 0.2–0.9 mm, distal margin entire, crenate, or deeply cleft or divided. |
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Staminate flowers | 5–20. |
25–35. |
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Pistillate flowers | ovary glabrous; styles 0.2–0.6 mm, 2-fid nearly 1/2 length. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
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Capsules | globose-ovoid, 1.5–2 × 1.5–3 mm, glabrous; columella 1.5–1.9 mm. |
depressed-ovoid to depressed-globose, 1.7–2.1 × 1.6–2.4 mm, glabrous; columella 1.2–1.8 mm. |
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Seeds | ashy white, ovoid, terete to bluntly subangled in cross section, 1.1–1.7 × 0.9–1.2 mm, smooth, with smooth brown line from top to bottom on adaxial side. |
white, ovoid-pyramidal, prominently 4-angled in cross section, 1.6–2 × 1–1.2 mm, smooth to slightly wrinkled. |
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Cyathia | solitary or in small, cymose clusters at distal nodes; peduncle 1–2 mm. |
solitary at distal nodes; peduncle 0.8–1.3 mm. |
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Euphorbia geyeri |
Euphorbia chaetocalyx |
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Distribution |
CO; IA; IL; IN; KS; LA; MI; MN; MO; MS; MT; ND; NE; NM; OK; SD; TX; UT; WI; WY; MB; n Mexico
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AZ; NM; TX; n Mexico
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Discussion | Varieties 2 (2 in the flora). The two varieties of Euphorbia geyeri have been distinguished in large part by the presence of conspicuous involucral gland appendages in var. geyeri and the lack of appendages in var. wheeleriana. The two varieties are recognized here, but the variation in the size and presence of involucral gland appendages in the closely related E. bombensis suggests that this might be a somewhat variable character in this group of species. Euphorbia geyeri is widespread throughout the central United States in sandy soils. Populations at the eastern edge of the range are often considered adventive (for example, sandy soils along railroad grades in Michigan). Euphorbia geyeri resembles E. glyptosperma (both being entirely glabrous), but that species has serrulate leaves (near the apex) and strongly angled, transverse-ridged seeds whereas E. geyeri has entire leaves and smooth, rounded seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia chaetocalyx is similar to E. fendleri but can generally be distinguished from that species by its narrow, acute leaves and ± erect stems. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. chaetocalyx from E. fendleri, but those characters appear highly variable and of little taxonomic utility. Some individuals from western Texas (Culberson and El Paso counties) and southern New Mexico appear intermediate with E. fendleri. The specific epithet of E. chaetocalyx refers to the bristly perianthlike segments that subtend the ovary, but these structures are found intermittently in both E. chaetocalyx and E. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 268. | FNA vol. 12, p. 263. | ||||||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||||||
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Subordinate taxa | ||||||||||
Synonyms | Chamaesyce geyeri | E. fendleri var. chaetocalyx, Chamaesyce chaetocalyx | ||||||||
Name authority | Engelmann: Boston J. Nat. Hist. 5: 260. (1845) | (Boissier) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | ||||||||
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