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greater Florida spurge

Hoover's sandmat, Hoover's spurge

Habit Herbs, perennial, with slender to moderately thickened rootstock. Herbs, annual, with slender to slightly thickened taproot.
Stems

erect, 20–65 cm.

prostrate, occasionally mat-forming, 4–10 cm, glabrous.

Leaves

petiole absent;

blade linear to linear-elliptic or linear-lanceolate, 30–105 × 2–4(–7) mm, chartaceous, base attenuate, rounded, or nearly truncate, apex acute;

only midvein evident.

opposite;

stipules distinct or connate at base, deeply laciniate into numerous subulate to filiform segments, 0.3–1.3 mm, glabrous;

petiole 0–0.5 mm, glabrous;

blade broadly ovate, broadly oblong or suborbiculate, 3–7 × 2–5.5 mm, base asymmetric, hemiamplexicaulous, margin sharply serrulate to denticulate, teeth whitish and occasionally setaelike at apex, apex obtuse, surfaces papillate, glabrous; 3-veined from base, usually only midvein conspicuous.

Involucre

campanulate or obconic, 1.6–3.3 × 1.8–3.1 mm, lobes ovate, 0.5–0.7 mm, ciliate, exceeded by glands;

glands yellow-green, oblong to trapezoidal, 0.5–0.8 × 0.8–1.2 mm, distal margins deeply erose.

campanulate, 1.2–1.5 × 1.5–2.2 mm, papillate;

glands 4, yellow to reddish, subcircular to oblong, 0.4–0.5 × 0.5–0.7 mm;

appendages white to pink, 0.3–0.9 × 0.7–1.6 mm, divided into 4–6 narrowly triangular, acute segments, these occasionally 2-fid, margins entire.

Staminate flowers

20.

25–35.

Pistillate flowers

gynophore exserted 2.9–5.3 mm, calyxlike lobes triangular, 0.3–0.7 mm;

styles connate 1/3–1/4 length, 1.1–1.7 mm.

ovary glabrous;

styles 1.8–2.6 mm, unbranched, filiform.

Capsules

oblate-ovoid, 4.6–5.5 × 8.9–9 mm, strongly 3-lobed;

columella 3.4–4.2 mm.

oblate, 1.5–1.7 × 1.8–2.2 mm, glabrous;

columella 1.2–1.4 mm.

Seeds

brown to blackish, depressed-globose, circular in cross section, 2.8–3 × 3.2–3.6 mm, smooth, base flattened, with punctiform depressions, apex flattened.

gray to light brown, ovoid, inconspicuously 4-angled in cross section, 1.2–1.6 × 1–1.1 mm, rugose.

Cyathial

arrangement: terminal pleiochasial branches 3–5, 3–16 cm, 3–5(–7) times 2-branched;

pleiochasial bracts linear-lanceolate or narrowly ovate, 19–55 × 4–12 mm, margins entire, apex acute or acuminate;

dichasial bracts usually ovate, lanceolate, or oblong, rarely deltate, 8–21 × 4–14 mm, margins entire, apex acute, acuminate, or rounded with mucronate tip;

axillary cymose branches 1–3(–6).

Cyathia

peduncle (except for that of 1st cyathium at base of pleiochasia) 2–5 mm (not exceeding dichasial bracts).

solitary at distal nodes;

peduncle 0.5 mm, stout.

Euphorbia floridana

Euphorbia hooveri

Phenology Flowering and fruiting spring–fall. Flowering and fruiting summer–fall.
Habitat Xeric pine-oak sandhills, pine scrub, sandy soils. Drying mudflats of vernal pools in grasslands and woodlands.
Elevation 20–80 m. (100–300 ft.) 20–200 m. (100–700 ft.)
Distribution
from FNA
AL; FL; GA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Euphorbia hooveri is federally listed as threatened; its populations are being affected severely by habitat loss and the invasion of exotic species. The species is endemic to vernal pools in six counties in the Central Valley of California. Molecular data show that E. hooveri is a hybrid species, closely related to E. albomarginata and E. serpens (Y. Yang and P. E. Berry 2011).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 314. FNA vol. 12, p. 271.
Parent taxa Euphorbiaceae > Euphorbia > sect. Nummulariopsis Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Galarhoeus floridanus Chamaesyce hooveri
Name authority Chapman: Fl. South. U.S., 401. (1860) L. C. Wheeler: Proc. Biol. Soc. Wash. 53: 9. (1940)
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