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Chiricahua Mountain sandmat

Roemer's spurge

Habit Herbs, annual, with slender taproot. Herbs, annual, with taproot.
Stems

erect, 15–60 cm, usually glabrous, rarely puberulent.

erect, occasionally decumbent at base, branched, 15–30 cm, glabrous.

Leaves

opposite;

stipules distinct, divided into 3–4 subulate-filiform divisions, 0.4–1.6 mm, usually glabrous, rarely puberulent;

petiole 0.5–2.5 mm, glabrous;

blade usually linear, rarely to narrowly elliptic, 10–40(–60) × 0.5–2.5 mm, base symmetric, attenuate, margins serrulate, often revolute, apex acute, surfaces usually glabrous, rarely puberulent; obscurely pinnately veined.

petiole 2–5 mm;

blade oblanceolate to obovate, 5–20 × 5–10 mm (larger leaves in distal portion of stem), base cuneate to attenuate, margins entire, apex rounded, surfaces glabrous;

venation pinnate.

Involucre

obconic, 1.7–2.4 × 1.5–2.1 mm, glabrous;

glands 4, greenish yellow to slightly pink, circular to oblong, 0.4–0.5 × 0.4–0.6 mm;

appendages white to pink, obovoid, circular, flabellate, or oblong, 0.8–2.9 × 1–2.8 mm, distal margin entire.

campanulate, 1.5–2 × 1–1.6 mm, glabrous;

glands 4, elliptic to trapezoidal, 0.4–0.6 × 0.7–1.2 mm;

horns slightly convergent, 0.1–0.3 mm.

Staminate flowers

25–35.

10–12.

Pistillate flowers

ovary glabrous;

styles 0.8–1.4 mm, 2-fid entire length.

ovary glabrous;

styles 0.6–1 mm, 2-fid.

Capsules

oblate, 2.2–2.5 × 2.7–3.1 mm, glabrous;

columella 1.8–2.1 mm.

subglobose, 2–2.5 × 2.4–3 mm, slightly lobed;

cocci rounded, smooth, glabrous;

columella 1.5–1.8 mm.

Seeds

light gray to light brown, ovoid, slightly 4-angled in cross section, 1.6–2 × 1.3–1.7 mm, with 2 or 3 well-developed transverse ridges.

brown, oblong-ovoid, 1.6–1.8 × 1.4–1.5 mm, with scattered, deep and broad pits;

caruncle conic, reniform, 0.3–0.4 × 0.4–0.6 mm.

Cyathia

solitary at nodes or in small, cymose clusters at branch tips;

peduncle 1.2–8.1 mm.

peduncle 0–0.9 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, 1–4 times 2-branched;

pleiochasial bracts ovate to oblong, similar in size to distal leaves;

dichasial bracts connate 1/2 length, reniform to semicircular, base truncate to perfoliate, margins entire, apex rounded to obtuse;

axillary cymose branches 0–3.

Euphorbia florida

Euphorbia roemeriana

Phenology Flowering and fruiting summer–late fall. Flowering and fruiting spring.
Habitat Sandy flats, gravelly washes, rocky hillsides, talus slopes, desert scrub, desert grasslands, mesquite woodlands, rarely oak woodlands. Rich calcareous soils, creek canyons.
Elevation 600–1300 m. (2000–4300 ft.) 100–300 m. (300–1000 ft.)
Distribution
from FNA
AZ; Mexico (Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Euphorbia florida is known in the flora area from Coconino County south to the Mexican border

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. S. Correll and M. C. Johnston (1970) suggested that Euphorbia roemeriana, which is restricted to the eastern part of the Edwards Plateau, is the southern counterpart of E. commutata, to which it is morphologically very similar. However, molecular phylogenetic analyses indicate that it is most closely related to E. austrotexana and E. longicruris (J. A. Peirson et al. 2014). Euphorbia roemeriana can be distinguished from E. commutata and several of the other small, annual species of subg. Esula by its consistently connate dichasial bracts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 267. FNA vol. 12, p. 308.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce florida Tithymalus roemerianus
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) Scheele: Linnaea 22: 151. (1849)
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