Euphorbia florida |
Euphorbia revoluta |
|
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Chiricahua Mountain sandmat |
revolute spurge, threadstem sandmat, threadstem spurge |
|
Habit | Herbs, annual, with slender taproot. | Herbs, annual, with slender taproot. |
Stems | erect, 15–60 cm, usually glabrous, rarely puberulent. |
erect, 5–25 cm, glabrous. |
Leaves | opposite; stipules distinct, divided into 3–4 subulate-filiform divisions, 0.4–1.6 mm, usually glabrous, rarely puberulent; petiole 0.5–2.5 mm, glabrous; blade usually linear, rarely to narrowly elliptic, 10–40(–60) × 0.5–2.5 mm, base symmetric, attenuate, margins serrulate, often revolute, apex acute, surfaces usually glabrous, rarely puberulent; obscurely pinnately veined. |
opposite; stipules distinct, subulate-filiform, entire, 0.5–0.9 mm, glabrous; petiole 0.6–1.1 mm, glabrous; blade linear, 6–27 × 0.6–1.2 mm, base symmetric, attenuate, margins entire, revolute, apex acute, sometimes mucronate, surfaces glabrous; only midvein conspicuous. |
Involucre | obconic, 1.7–2.4 × 1.5–2.1 mm, glabrous; glands 4, greenish yellow to slightly pink, circular to oblong, 0.4–0.5 × 0.4–0.6 mm; appendages white to pink, obovoid, circular, flabellate, or oblong, 0.8–2.9 × 1–2.8 mm, distal margin entire. |
obconic to campanulate, 0.7–0.9 × 0.5–0.7 mm, glabrous; glands 4, pink to dark purple, nearly circular to oblong or reniform, 0.1 × 0.1–0.2 mm; appendages white, oblong, nearly circular, ovate, deltate, or forming thin margin around gland, rarely absent, (0–)0.1–0.2 × (0–)0.1–0.3 mm, distal margin entire. |
Staminate flowers | 25–35. |
5–10. |
Pistillate flowers | ovary glabrous; styles 0.8–1.4 mm, 2-fid entire length. |
ovary glabrous; styles 0.2–0.3 mm, unbranched. |
Capsules | oblate, 2.2–2.5 × 2.7–3.1 mm, glabrous; columella 1.8–2.1 mm. |
ovoid to subglobose, 1.5–1.8 × 1.6–1.8 mm, glabrous; columella 1.1–1.5 mm. |
Seeds | light gray to light brown, ovoid, slightly 4-angled in cross section, 1.6–2 × 1.3–1.7 mm, with 2 or 3 well-developed transverse ridges. |
whitish, brick red, light gray, or light brown, narrowly to broadly ovoid, 4-angled in cross section, 0.9–1.4 × 0.7–1 mm, nearly smooth, rugulose, with faint transverse ridges, or with 2–3 well-defined transverse ridges separated by shallow depressions. |
Cyathia | solitary at nodes or in small, cymose clusters at branch tips; peduncle 1.2–8.1 mm. |
solitary at distal nodes; peduncles absent or to 1.5 mm. |
Euphorbia florida |
Euphorbia revoluta |
|
Phenology | Flowering and fruiting summer–late fall. | Flowering and fruiting summer–late fall. |
Habitat | Sandy flats, gravelly washes, rocky hillsides, talus slopes, desert scrub, desert grasslands, mesquite woodlands, rarely oak woodlands. | Desert scrub, sagebrush scrub, juniper woodlands, Joshua tree-pinyon pine woodlands, oak woodlands, grasslands, chaparral, pine-oak forests. |
Elevation | 600–1300 m. (2000–4300 ft.) | 600–2500 m. (2000–8200 ft.) |
Distribution |
AZ; Mexico (Sinaloa, Sonora)
|
AZ; CA; CO; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Sonora, Zacatecas)
|
Discussion | Euphorbia florida is known in the flora area from Coconino County south to the Mexican border (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia revoluta is a distinctive, easily recognizable species by virtue of the combination of unbranched styles and linear leaves with symmetric bases. The species ranges through northern Mexico into the southwestern United States and is composed of three well-marked, geographically distinct races that can be distinguished by their seeds. The first of these races occurs primarily in pine-oak forest of the Sierra Madre Occidental of Mexico (Chihuahua and Sonora) and barely enters the flora area in the Huachuca Mountains of southeastern Arizona; the seeds are brick red and nearly smooth. The second race is widespread in the Chihuahuan Desert from northern Mexico to southeastern New Mexico and southwestern Texas; its seeds are whitish to light gray and possess two or three well-defined transverse ridges separated by shallow depressions. The third race corresponds to the type collection and is characterized by grayish white to light brown seeds that are rugulose or with faint transverse ridges; it occurs in northwestern Mexico (Baja California and Sonora) and throughout the arid southwestern United States. It is probable that further study will justify the taxonomic segregation of these races as either distinct species or subspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 267. | FNA vol. 12, p. 286. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce florida | Chamaesyce revoluta |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) |
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