Euphorbia florida |
Euphorbia nutans |
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Chiricahua Mountain sandmat |
eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, annual, with taproot. |
Stems | erect, 15–60 cm, usually glabrous, rarely puberulent. |
usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
Leaves | opposite; stipules distinct, divided into 3–4 subulate-filiform divisions, 0.4–1.6 mm, usually glabrous, rarely puberulent; petiole 0.5–2.5 mm, glabrous; blade usually linear, rarely to narrowly elliptic, 10–40(–60) × 0.5–2.5 mm, base symmetric, attenuate, margins serrulate, often revolute, apex acute, surfaces usually glabrous, rarely puberulent; obscurely pinnately veined. |
opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
Involucre | obconic, 1.7–2.4 × 1.5–2.1 mm, glabrous; glands 4, greenish yellow to slightly pink, circular to oblong, 0.4–0.5 × 0.4–0.6 mm; appendages white to pink, obovoid, circular, flabellate, or oblong, 0.8–2.9 × 1–2.8 mm, distal margin entire. |
narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
Staminate flowers | 25–35. |
5–28. |
Pistillate flowers | ovary glabrous; styles 0.8–1.4 mm, 2-fid entire length. |
ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
Capsules | oblate, 2.2–2.5 × 2.7–3.1 mm, glabrous; columella 1.8–2.1 mm. |
ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
Seeds | light gray to light brown, ovoid, slightly 4-angled in cross section, 1.6–2 × 1.3–1.7 mm, with 2 or 3 well-developed transverse ridges. |
dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
Cyathia | solitary at nodes or in small, cymose clusters at branch tips; peduncle 1.2–8.1 mm. |
solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
2n | = 12, 14, 22. |
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Euphorbia florida |
Euphorbia nutans |
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Phenology | Flowering and fruiting summer–late fall. | Flowering and fruiting spring–early fall. |
Habitat | Sandy flats, gravelly washes, rocky hillsides, talus slopes, desert scrub, desert grasslands, mesquite woodlands, rarely oak woodlands. | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. |
Elevation | 600–1300 m. (2000–4300 ft.) | 0–1600 m. (0–5200 ft.) |
Distribution |
AZ; Mexico (Sinaloa, Sonora)
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AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
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Discussion | Euphorbia florida is known in the flora area from Coconino County south to the Mexican border (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 267. | FNA vol. 12, p. 278. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce florida | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Lagasca: Gen. Sp. Pl., 17. (1816) |
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