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Chiricahua Mountain sandmat

Heller's spurge

Habit Herbs, annual, with slender taproot. Herbs, annual, with taproot.
Stems

erect, 15–60 cm, usually glabrous, rarely puberulent.

ascending, branched proximally, 15–30 cm, glabrous.

Leaves

opposite;

stipules distinct, divided into 3–4 subulate-filiform divisions, 0.4–1.6 mm, usually glabrous, rarely puberulent;

petiole 0.5–2.5 mm, glabrous;

blade usually linear, rarely to narrowly elliptic, 10–40(–60) × 0.5–2.5 mm, base symmetric, attenuate, margins serrulate, often revolute, apex acute, surfaces usually glabrous, rarely puberulent; obscurely pinnately veined.

petiole usually 1–3 mm, absent distally;

blade spatulate, 6–15 × 3–5 mm, base broadly attenuate, margins entire, apex usually obtuse to rounded, sometimes retuse, surfaces glabrous;

venation pinnate, midvein prominent.

Involucre

obconic, 1.7–2.4 × 1.5–2.1 mm, glabrous;

glands 4, greenish yellow to slightly pink, circular to oblong, 0.4–0.5 × 0.4–0.6 mm;

appendages white to pink, obovoid, circular, flabellate, or oblong, 0.8–2.9 × 1–2.8 mm, distal margin entire.

campanulate, 0.9–1.2 × 0.7–1 mm, glabrous;

glands 4, elliptic, 0.1–0.2 × 0.3–0.4 mm;

horns slightly divergent, 0.1–0.2 mm.

Staminate flowers

25–35.

8–10.

Pistillate flowers

ovary glabrous;

styles 0.8–1.4 mm, 2-fid entire length.

ovary glabrous;

styles 0.4–0.5 mm, 2-fid.

Capsules

oblate, 2.2–2.5 × 2.7–3.1 mm, glabrous;

columella 1.8–2.1 mm.

depressed-globose, 2.1–2.5 × 2.5–3 mm, 3-lobed;

cocci flattened, smooth, glabrous;

columella 1.2–1.7 mm.

Seeds

light gray to light brown, ovoid, slightly 4-angled in cross section, 1.6–2 × 1.3–1.7 mm, with 2 or 3 well-developed transverse ridges.

white to light gray, ovoid, 1.4–1.6 × 1–1.2 mm, smooth;

caruncle 2-lobed, thin, 0.4 × 0.7 mm.

Cyathia

solitary at nodes or in small, cymose clusters at branch tips;

peduncle 1.2–8.1 mm.

peduncle 0.2–0.4 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, each 1–5 times 2-branched;

pleiochasial bracts oblong, similar in size to distal leaves;

dichasial bracts distinct, orbiculate-ovate to nearly reniform, subpandurate, base broadly cuneate to truncate, margins entire, apex obtuse, mucronate;

axillary cymose branches 0–5.

Euphorbia florida

Euphorbia helleri

Phenology Flowering and fruiting summer–late fall. Flowering and fruiting late winter–spring.
Habitat Sandy flats, gravelly washes, rocky hillsides, talus slopes, desert scrub, desert grasslands, mesquite woodlands, rarely oak woodlands. Forests, stream banks, roadsides, shaded areas with sandy, calcareous soils.
Elevation 600–1300 m. (2000–4300 ft.) 0–50 m. (0–200 ft.)
Distribution
from FNA
AZ; Mexico (Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; TX; Mexico (Nuevo León)
[BONAP county map]
Discussion

Euphorbia florida is known in the flora area from Coconino County south to the Mexican border

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The smooth, white to light gray seeds of Euphorbia helleri easily distinguish it from other annual members of subg. Esula in North America. Collections of E. helleri have been made near Brownsville, Texas, and thus it is possible that the species occurs in northern Tamaulipas, Mexico. The Louisiana record (Webster Parish) likely represents introduced plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 267. FNA vol. 12, p. 303.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce florida Tithymalus helleri
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) Millspaugh: Bot. Gaz. 26: 268, fig. [p. 270]. (1898)
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