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Fendler's sandmat
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sawtooth sandmat
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| Herbs, perennial, with woody, thickened taproot. |
Herbs, annual, with slender taproot. |
usually prostrate, decumbent, or ascending, very rarely erect, often densely clustered from top of woody crown, 5–12 cm, glabrous. |
prostrate or ascending, 5–20 cm, usually pilose to villous, rarely glabrate. |
usually opposite, rarely whorled; stipules distinct, narrowly linear, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade usually orbiculate to ovate, rarely almost lanceolate, 3–8 × 2.5–7 mm, base slightly asymmetric, slightly cordate to rounded or obtuse, margins entire, apex rounded to obtuse, surfaces glabrous; obscurely 3–5-veined from base, only midvein conspicuous. |
opposite; stipules usually distinct, rarely connate at base, triangular or laciniate into subulate segments, 1–1.8 mm, glabrous; petiole 0.3–0.8 mm, glabrous or villous; blade oblong, ovate, or elliptic, sometimes falcate, 3–11 × 2–5 mm, base asymmetric, rounded to hemicordate, margins sharply serrate to serrulate, usually with conspicuous teeth at base of leaf, apex usually obtuse, rarely acute, surfaces frequently with red blotch in center, not papillate, sparsely pilose to glabrate; only midvein conspicuous. |
campanulate to turbinate or broadly cupulate, 1.1–1.7 × 1.2–1.8 mm, glabrous; glands 4, yellow-green to reddish, elliptic to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages absent or white, rarely pink, often unequal, lunate to flabellate or sometimes forming crenate margin along gland, (0–)0.1–0.6 × (0–)0.5–1.5 mm, distal margin entire or toothed. |
obconic, 0.8–1.1 × 0.8–1 mm, glabrous; glands 4, greenish yellow, usually reniform to elliptic, rarely circular, 0.1 × 0.1–0.2 mm; appendages usually white, rarely light pink, orbiculate, 0.1–0.3 × 0.2–0.4 mm, distal margin entire or crenulate. |
25–35. |
7–15. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
depressed-globose, 2–2.4 × 2.2–2.5 mm, glabrous; columella 1.7–2.1 mm. |
oblate, cocci not elongated nor terminating in empty portion, 2–2.6 × 3.2–3.7 mm, glabrous; columella 1.7–2.1 mm. |
white, ovoid-pyramidal, prominently 4-angled in cross section, 1.7–2 × 1–1.2 mm, smooth to slightly wrinkled. |
white to light brown, broadly ellipsoid to ovoid, 3–4-angled in cross section, 1.5–1.8 × 1.1–1.3(–1.5) mm, smooth to minutely rugulose or with scattered small depressions. |
solitary at distal nodes; peduncle 0.7–1.2 mm. |
solitary at distal nodes; peduncle 0.4–1.8(–2.3) mm. |
= 28. |
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| Flowering and fruiting spring–fall. |
Flowering and fruiting spring–fall. |
| Mountain slopes, desert scrub, pinyon-juniper woodlands, hills, canyons, grasslands, washes, roadsides, dry crevices in limestone, often in gravel and sand. |
Desert scrub, with creosote bush, grasslands with mesquite and yucca, rarely in ponderosa pine woodlands, often sandy substrates. |
| 500–2600 m. (1600–8500 ft.) |
300–1900 m. (1000–6200 ft.) |
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AZ; CA; CO; KS; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Coahuila)
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AZ; NM; TX; Mexico
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Euphorbia fendleri is similar to E. chaetocalyx and may sometimes be confused with that species. Its prostrate to decumbent or ascending stems and small, ovate to orbiculate leaves distinguish it from E. chaetocalyx. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. fendleri from E. chaetocalyx, but those characters appear highly variable and of little taxonomic utility; somewhat intermediate individuals occur in western Arizona, New Mexico, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In Mexico, Euphorbia serrula is found from Chihuahua and Coahuila south to Puebla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 267. |
FNA vol. 12, p. 288. |
| Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| Chamaesyce fendleri |
Chamaesyce serrula |
| Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 175. (1857) |
Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) |
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