Euphorbia fendleri |
Euphorbia exserta |
|
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Fendler's sandmat |
coastal sand spurge, maroon or purple sand spurge |
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Habit | Herbs, perennial, with woody, thickened taproot. | Herbs, perennial, with spreading rootstock. |
Stems | usually prostrate, decumbent, or ascending, very rarely erect, often densely clustered from top of woody crown, 5–12 cm, glabrous. |
erect or ascending, unbranched or branched, solitary, few, or occasionally densely clumped, previous year's dead stems not persistent, 20–33 cm, usually glabrous, rarely sparsely villous, glaucescent. |
Leaves | usually opposite, rarely whorled; stipules distinct, narrowly linear, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade usually orbiculate to ovate, rarely almost lanceolate, 3–8 × 2.5–7 mm, base slightly asymmetric, slightly cordate to rounded or obtuse, margins entire, apex rounded to obtuse, surfaces glabrous; obscurely 3–5-veined from base, only midvein conspicuous. |
alternate; stipules less than 0.1 mm; petiole (0–)1–3 mm, glabrous; blade linear or linear-elliptic to obovate or orbiculate, proximal greatly reduced, scalelike, 15–30 × 1–20 mm, base cuneate to rounded, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous, glaucescent; venation often obscure on narrow leaves, midvein conspicuous. |
Involucre | campanulate to turbinate or broadly cupulate, 1.1–1.7 × 1.2–1.8 mm, glabrous; glands 4, yellow-green to reddish, elliptic to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages absent or white, rarely pink, often unequal, lunate to flabellate or sometimes forming crenate margin along gland, (0–)0.1–0.6 × (0–)0.5–1.5 mm, distal margin entire or toothed. |
usually dark red, campanulate, 1.3–1.6 × 1.4–2.1 mm, glabrous; glands 5, usually dark red, rarely greenish red, elliptic reniform, thickened, 0.3–0.5 × 0.8 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
Staminate flowers | 25–35. |
20–25. |
Pistillate flowers | ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
Capsules | depressed-globose, 2–2.4 × 2.2–2.5 mm, glabrous; columella 1.7–2.1 mm. |
depressed-globose, 1.8–2.5 × 3.6–4.4 mm, glabrous; columella 1.9–2.4 mm. |
Seeds | white, ovoid-pyramidal, prominently 4-angled in cross section, 1.7–2 × 1–1.2 mm, smooth to slightly wrinkled. |
ashy white, ovoid, 2.1 × 1.3 mm, angled with 5 blunt longitudinal ridges, with shallow and irregular pits; caruncle absent. |
Cyathia | solitary at distal nodes; peduncle 0.7–1.2 mm. |
usually in terminal dichasia, sometimes pleiochasia; peduncle 6–33 mm, filiform, glabrous. |
2n | = 28. |
|
Euphorbia fendleri |
Euphorbia exserta |
|
Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting spring–summer. |
Habitat | Mountain slopes, desert scrub, pinyon-juniper woodlands, hills, canyons, grasslands, washes, roadsides, dry crevices in limestone, often in gravel and sand. | Xeric to dry pine-oak scrub of sand hills, pine-oak woodlands, pine-oak savannas. |
Elevation | 500–2600 m. (1600–8500 ft.) | 0–150 m. (0–500 ft.) |
Distribution |
AZ; CA; CO; KS; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Coahuila)
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FL; GA; NC; SC; VA
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Discussion | Euphorbia fendleri is similar to E. chaetocalyx and may sometimes be confused with that species. Its prostrate to decumbent or ascending stems and small, ovate to orbiculate leaves distinguish it from E. chaetocalyx. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. fendleri from E. chaetocalyx, but those characters appear highly variable and of little taxonomic utility; somewhat intermediate individuals occur in western Arizona, New Mexico, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
K. R. Park (1998) recognized both Euphorbia exserta and E. gracilior as distinct species, with the former known only from the holotype. However, this treatment follows M. J. Huft (1997) and treats E. gracilior as a synonym of E. exserta. Although the type of E. exserta is unusual in having greenish red (versus dark red) cyathia and glands, as well as small gland appendages, it is otherwise typical of the species as a whole, including features such as the upright habit, reddish coloration, scalelike proximal leaves, and filiform peduncles to 30 mm that are also common to plants formerly treated as E. gracilior. Euphorbia gracilis Elliott, which has sometimes been applied to E. exserta, is an illegitimate name (a later homonym of E. gracilis Loiseleur-Deslongchamps) and pertains here. The upright habit and usually dark red cyathia and glands distinguish Euphorbia exserta from the otherwise similar E. ipecacuanhae, while the glaucescent vegetative parts and smaller gland appendages readily separate E. exserta from the similar E. curtisii. Euphorbia exserta is found on the Gulf and Atlantic coastal plains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 267. | FNA vol. 12, p. 246. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce fendleri | Tithymalopsis exserta, E. gracilior, T. gracilis |
Name authority | Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 175. (1857) | (Small) Coker: Pl. Life Hartsville, 88. (1912) |
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