Euphorbia exstipulata |
Euphorbia peplus |
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square-seed spurge, squareseed or Clark Mountain spurge |
petty spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, annual, with taproot. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
erect, unbranched or branched, 5–30 cm, glabrous. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
petiole 1–10 mm; blade obovate, oblong, or suborbiculate, 5–25 × 4–15 mm, base attenuate or cuneate, margins entire, apex obtuse, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
cupulate to slightly turbinate, 0.6–1.1 × 0.7–0.9 mm, glabrous; glands 4, elliptic, 0.2–0.3 × 0.5–0.6 mm; horns slightly convergent to divergent, 0.4–0.6 mm. |
Staminate flowers | 10–12. |
10–15. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.1–0.2 mm, 2-fid. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
subglobose, 1.3–2 × 1.5–2.2 mm, slightly 3-lobed; cocci rounded, with 2 low longitudinal wings, smooth, glabrous; columella 1.1–1.3 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
whitish or grayish, subovoid, 1–1.6 × 0.6–1 mm, abaxial faces regularly large-pitted (appearing almost alveolate), adaxial faces longitudinally sulcate; caruncle deciduous, conic, 0.2–0.5 × 0.2–0.7 mm. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
arrangement: terminal pleiochasial branches 3–5, usually 2–8 times 2-branched, 1st branching level rarely 3-furcate; pleiochasial bracts similar in shape to and slightly larger than distal leaves; dichasial bracts distinct, ovate to obovate, base obtuse, margins entire, apex obtuse, usually mucronate; axillary cymose branches 0–3. |
Cyathia | peduncle 1–1.9 mm. |
peduncle 0.4–1.1 mm. |
Euphorbia exstipulata |
Euphorbia peplus |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting year-round. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Edges of gardens, weedy flower beds, roadsides, waste places, open ground near forests. |
Elevation | 800–2000 m. (2600–6600 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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AL; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; TX; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe; w Asia [Introduced in North America; introduced also in Mexico, West Indies, Bermuda, Central America, South America, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
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Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 307. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | Esula peplus, Galarhoeus peplus, Tithymalus peplus |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Linnaeus: Sp. Pl. 1: 456. (1753) |
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