Euphorbia exstipulata |
Euphorbia myrsinites |
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square-seed spurge, squareseed or Clark Mountain spurge |
broad leaf glaucous spurge, donkey tail, myrtle or creeping or blue spurge, myrtle spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, usually perennial, occasionally biennial, with taproot. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
erect or semiprostrate, unbranched or branched, 15–40 cm, succulent, glabrous. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
petiole 0–2 mm; blade obovate, obovate-oblong, lanceolate, orbiculate, or suborbiculate, 2–30 × 3–17 mm, fleshy, base truncate or attenuate, margins entire or finely denticulate, apex acute to obtuse, cuspidate or strongly mucronate, surfaces glabrous; venation and midvein inconspicuous. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
campanulate, 2.4–2.6 × 2.3–2.5 mm, glabrous; glands 4, trapezoidal, 1–1.5 × 1.5–2.5 mm; horns divergent, thick, tips rounded, dilated, 0.5–0.9 mm. |
Staminate flowers | 10–12. |
6–12. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 2.5–2.8 mm, usually unbranched. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
subglobose, 5–7 × 5–6 mm, unlobed; cocci rounded to subangular, smooth, glabrous; columella 4.5–5 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
brownish to grayish, oblong, 2.8–4.5 × 2–3.2 mm, vermiculate-rugose; caruncle substipitate, trapezoidal or mushroom-shaped, 1.3–1.5 × 0.6–0.8 mm. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
arrangement: terminal pleiochasial branches 2–12, each 1–2 times 2-branched; pleiochasial bracts similar in shape and size to distal leaves; dichasial bracts distinct, suborbiculate or reniform, base truncate, margins entire or minutely denticulate, apex obtuse, mucronulate; axillary cymose branches 0–4. |
Cyathia | peduncle 1–1.9 mm. |
peduncle 0.5–1 mm. |
Euphorbia exstipulata |
Euphorbia myrsinites |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–summer. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Scrub oak communities, open ground near forests, shrub-steppes. |
Elevation | 800–2000 m. (2600–6600 ft.) | 0–2400 m. (0–7900 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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CA; CO; ID; MT; NM; OR; UT; WA; WY; BC; s Europe; w Asia [Introduced in North America]
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Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia myrsinites is cultivated in much of the flora area, where it can tolerate cold winters. In some areas, it can locally escape from cultivation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 305. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | Tithymalus myrsinites |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Linnaeus: Sp. Pl. 1: 461. (1753) |
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