Euphorbia exstipulata |
Euphorbia lata |
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square-seed spurge, squareseed or Clark Mountain spurge |
broadleaf spurge, hoary sandmat |
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Habit | Herbs, annual, with slender taproot. | Herbs, perennial, with moderately thickened to robust rootstock. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
ascending to erect, or prostrate, 10–25 cm, strigose to short-sericeous or ± villous. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
opposite; stipules distinct, filiform, 0.8–1.3 mm, strigose to short-sericeous or ± villous; petiole 0.5–2 mm, densely strigose to short-sericeous or ± villous; blade narrowly to broadly ovate-deltate, older ones often falcate, 4–12 × 3–7 mm, base asymmetric, obliquely rounded to obtuse, noticeably wider on one side, margins entire, often ± revolute, apex broadly acute, surfaces strigose to short-sericeous or ± villous; obscurely 3–5-veined from base, midvein prominent abaxially. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
broadly campanulate, 2–2.5 × 2.2–2.6, strigose; glands 4, greenish, oblong to semilunate, 0.2–0.7 × 0.6–1 mm; appendages rudimentary or white, forming narrow band, (0–)0.1–0.2 × (0–)0.6–1 mm, distal margin entire or crenate. |
Staminate flowers | 10–12. |
25–35. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary densely strigose to short-sericeous or ± villous; styles dark purplish, 0.8–1.2 mm, 2-fid 1/2 to nearly entire length. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
ovoid, 1.9–2.3 × 2–2.4 mm, strigose to short-sericeous or ± villous; columella 1.7–2.2 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
whitish, oblong, 4-angled in cross section, faces concave, 1.5–1.8(–2) × 0.6–0.9 mm, smooth. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
|
Cyathia | peduncle 1–1.9 mm. |
solitary at distal nodes; peduncle 1–3 mm. |
2n | = 28, 56. |
|
Euphorbia exstipulata |
Euphorbia lata |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–fall. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Mountain slopes, canyons, basins, rocky prairies, roadsides, disturbed sites, usually in calcareous soils, sometimes in igneous-derived, sandy or rocky soils. |
Elevation | 800–2000 m. (2600–6600 ft.) | 600–2200 m. (2000–7200 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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CO; KS; NM; OK; TX; Mexico (Chihuahua, Coahuila)
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Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 275. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | E. dilatata, Chamaesyce lata, E. rinconis |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) |
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