Euphorbia exstipulata |
Euphorbia innocua |
|
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square-seed spurge, squareseed or Clark Mountain spurge |
velvet spurge |
|
Habit | Herbs, annual, with slender taproot. | Herbs, perennial, with moderately to strongly thickened rootstock. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
prostrate to decumbent or ascending, branched (often near base), 7–45 cm, densely pilose. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
alternate; stipules to 0.1 mm; petiole (0.7–)1.1–3.5 mm, pilose; blade ovate to orbiculate, 4.6–17(–25) × 4.5–15(–19) mm, base cordate, margins entire, apex rounded to obtuse, surfaces densely pilose; venation obscure, usually only midvein conspicuous. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
campanulate, 1–1.3 × 1.2–1.4 mm, pilose; glands 4, yellow to green, elliptic, 0.2–0.3 × 0.5–0.6 mm; appendages green, elliptic, 0.4–0.5 × 0.5–0.9 mm, entire or crenulate, ciliate. |
Staminate flowers | 10–12. |
5–10. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary pilose; styles 0.4–0.7 mm, 2-fid 1/2 length. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
depressed-ovoid, 2–2.5 × 2.7–3.3 mm, pilose; columella 1.6–2.1 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
gray to brown, ovoid, 1.5–1.7 × 1.2–1.3 mm, rugose with whitish ridges; caruncle absent. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
|
Cyathia | peduncle 1–1.9 mm. |
in terminal dichasia (often weakly defined); peduncle 1–2.7 mm, densely pilose. |
Euphorbia exstipulata |
Euphorbia innocua |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting early winter–late spring. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Sandy soils or dunes, grasslands, pastures. |
Elevation | 800–2000 m. (2600–6600 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
|
TX |
Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia innocua is restricted to south coastal Texas in Aransas, Calhoun, Kenedy, Kleberg, Nueces, Refugio, San Patricio, and Willacy counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 247. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | L. C. Wheeler: Contr. Gray Herb. 127: 62, plate 3, fig. D. (1939) |
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