Euphorbia exstipulata |
Euphorbia georgiana |
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square-seed spurge, squareseed or Clark Mountain spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, annual, with taproot. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
erect, often branched near base, 10–18 cm, glabrous. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
petiole absent; blade oblanceolate, 5–12 × 3–5 mm, base attenuate, margins entire, apex rounded, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
infundibular, 1–1.1 × 0.4–0.5 mm, glabrous; glands 4, crescent-shaped; 0.3–0.4 × 0.4–0.5 mm; horns divergent, 0.3–0.5 mm. |
Staminate flowers | 10–12. |
5–10. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.5 mm, 2-fid. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
depressed-ovoid, 2.2–2.4 × 3.2–3.4 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 2–2.1 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
gray, ovoid, 1.6–1.7 × 1.4–1.6 mm, with deep, rounded pits irregularly scattered over entire surface; caruncle reniform, subconic, 0.5–0.6 × 0.6–0.8 mm. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched; pleiochasial bracts rotund-obovate, wider than distal leaves; dichasial bracts distinct, not imbricate, broadly deltate to subreniform, base truncate to emarginate, margins entire, apex rounded to bluntly acuminate; axillary cymose branches 0–1. |
Cyathia | peduncle 1–1.9 mm. |
peduncle 0.3–0.5 mm. |
Euphorbia exstipulata |
Euphorbia georgiana |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Granite outcrops. |
Elevation | 800–2000 m. (2600–6600 ft.) | 100–200 m. (300–700 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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GA |
Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia georgiana is restricted to granitic outcrops; it is known from Oglethorpe and Wilkes counties (M. H. Mayfield 2013). It is similar to E. austrotexana but has larger seeds that are much more deeply pitted and leaves that are oblanceolate instead of linear-oblanceolate to linear. It is also quite distinct from the more robust, biennial or occasionally annual E. commutata, the only other closely-related species that occurs in the area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 303. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Mayfield: J. Bot. Res. Inst. Texas 7: 639, fig. 2[row 2, left]. (2013) |
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