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square-seed spurge, squareseed or Clark Mountain spurge
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Fendler's sandmat
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| Herbs, annual, with slender taproot. |
Herbs, perennial, with woody, thickened taproot. |
erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
usually prostrate, decumbent, or ascending, very rarely erect, often densely clustered from top of woody crown, 5–12 cm, glabrous. |
opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
usually opposite, rarely whorled; stipules distinct, narrowly linear, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade usually orbiculate to ovate, rarely almost lanceolate, 3–8 × 2.5–7 mm, base slightly asymmetric, slightly cordate to rounded or obtuse, margins entire, apex rounded to obtuse, surfaces glabrous; obscurely 3–5-veined from base, only midvein conspicuous. |
turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
campanulate to turbinate or broadly cupulate, 1.1–1.7 × 1.2–1.8 mm, glabrous; glands 4, yellow-green to reddish, elliptic to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages absent or white, rarely pink, often unequal, lunate to flabellate or sometimes forming crenate margin along gland, (0–)0.1–0.6 × (0–)0.5–1.5 mm, distal margin entire or toothed. |
10–12. |
25–35. |
ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
depressed-globose, 2–2.4 × 2.2–2.5 mm, glabrous; columella 1.7–2.1 mm. |
white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
white, ovoid-pyramidal, prominently 4-angled in cross section, 1.7–2 × 1–1.2 mm, smooth to slightly wrinkled. |
arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
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peduncle 1–1.9 mm. |
solitary at distal nodes; peduncle 0.7–1.2 mm. |
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= 28. |
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| Flowering and fruiting summer–fall. |
Flowering and fruiting spring–fall. |
| Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. |
Mountain slopes, desert scrub, pinyon-juniper woodlands, hills, canyons, grasslands, washes, roadsides, dry crevices in limestone, often in gravel and sand. |
| 800–2000 m. (2600–6600 ft.) |
500–2600 m. (1600–8500 ft.) |
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AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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AZ; CA; CO; KS; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Coahuila)
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Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia fendleri is similar to E. chaetocalyx and may sometimes be confused with that species. Its prostrate to decumbent or ascending stems and small, ovate to orbiculate leaves distinguish it from E. chaetocalyx. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. fendleri from E. chaetocalyx, but those characters appear highly variable and of little taxonomic utility; somewhat intermediate individuals occur in western Arizona, New Mexico, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 322. |
FNA vol. 12, p. 267. |
| Euphorbiaceae > Euphorbia > sect. Poinsettia |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| E. exstipulata var. lata |
Chamaesyce fendleri |
| Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 175. (1857) |
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