Euphorbia exstipulata |
Euphorbia conferta |
|
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square-seed spurge, squareseed or Clark Mountain spurge |
Everglade key sandmat |
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Habit | Herbs, annual, with slender taproot. | Herbs, annual, with taproot. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
prostrate, ascending, to erect, 10–45 cm, pilose. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
opposite; stipules distinct, subulate, 1–2 mm, pilose; petiole 0.9–1.5 mm, pilose; blade oblanceolate to elliptic, 3–10 × 1–4 mm, base asymmetric, obtuse to hemicordate, margins coarsely serrate, sometimes revolute, apex rounded to acute, abaxial surface sparsely hispidulous to strigillose (densely so on young leaves), adaxial surface glabrous; 3-veined from base but only midvein conspicuous. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
campanulate, 1.1–1.4 × 0.5–0.7 mm, pilose; glands 4, red, narrowly reniform, 0.1 × 0.2–0.4 mm; appendages pink to red, larger 2 petal-like, smaller 2 elliptic, 0.4–1 × 0.3–0.5 mm, distal margin erose. |
Staminate flowers | 10–12. |
5–8. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary pilose; styles 0.4–0.8 mm, 2-fid at apex. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
conic or truncate-ovoid, 1–1.2 mm diam., pilose; columella 0.9–1.1 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
orange-brown, glaucous, oblong, 4-angled in cross section, 0.7–0.8 × 0.4–0.5 mm, slightly rugose and with 3–4 transverse ridges. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
|
Cyathia | peduncle 1–1.9 mm. |
solitary at distal nodes (appearing clustered at points of new growth); peduncle 0.4–0.6 mm. |
Euphorbia exstipulata |
Euphorbia conferta |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting year-round. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Sandy, disturbed, wet areas, often roadsides. |
Elevation | 800–2000 m. (2600–6600 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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FL |
Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia conferta is known only from Broward, Miami-Dade, and Monroe counties in southern Florida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 264. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | Chamaesyce conferta |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | (Small) B. E. Smith: J. Elisha Mitchell Sci. Soc. 62: 82. (1946) |
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